The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae. These centipedes are found almost worldwide.
Geophilus is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with Brachygeophilus. This genus is mostly Holarctic and characterized by a claw-shaped ultimate pretarsus, complete or nearly complete coxo-pleural sutures at the prosternum, and incomplete chitin-lines. Centipedes in this genus feature slightly elongate heads and labral intermediate parts with tubercles, the forcipules are usually poorly elongate with a single small tubercle at the base of each tarsungulum, and the anterior trunk metasternites usually have an anterior medial socket and a posterior transversally elongate pore-field. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as Geophilus electricus.
Geophilus persephones is a species of soil centipede in the family Geophilidae discovered in 1999. This species is named after Persephone, the queen of the underworld in Greek mythology, and found in caves in the Gouffre de la Pierre Saint-Martin. It has elongated antennae and legs as well as abundant sensory setae, and like other geophilomorhps it lacks sight, has a flattened trunk, and is well adapted to underground life. This species was the first troglomorphic geophilomorph ever discovered and one of the only two in existence along with Geophilus hadesi. Known from a single male specimen measuring 16.2 mm in length, this species has only 29 pairs of legs, one of only two species in the Geophilidae family to have so few leg pairs.
Geophilus electricus is a species of soil centipede in the family Geophilidae found across temperate Europe, especially Britain and Ireland. Originally discovered and named Scolopendra electrica in 1758, it was later moved to its own genus, Geophilus. As the first species described in this genus, it is considered by some to be the type species. It is yellowish-orange in color and can grow up to 45 millimeters, with 55 to 75 leg pairs, a distinct carpophagus fossae on the anterior sternites, and a distinctive arrangement of the coxal pores of the last leg pair. Like many geophilomorphans, G. electricus has been known to glow in the dark, most likely as a defensive maneuver.
Geophilus carpophagus is a species of soil centipede in the family Geophilidae, widely considered to be a type species of the genus Geophilus. It grows up to 60 millimeters in length, with an orange/tan body bearing a distinctive purplish marbled pattern. Males of this species have 51 to 55 pairs of legs; females have 53 to 57.
Geophilus arenarius is a species of soil centipede in the family Geophilidae found in northwest Africa, specifically near Annaba, Algeria. This species is frequently misidentified with G. electricus, and as part of the carpophagus species-complex, is closely related to both G. carpophagus and G. easoni, though it differs mainly by lacking a transverse suture on the head and peculiar integumental features (carpophagus-structures) along the trunk, as well as having relatively stouter antennae and forcipular coxosternite. G. arenarius is distinctly smaller at full growth than G. carpophagus, with usually blunter and more sclerotised tubercles lining the intermediate part of the labrum and a minute denticle at the basis of the forcipular tarsungula. It has fewer bristles lining the lateral parts of the labrum than G. easoni as well as a generally higher number of legs and a more greyish coloured trunk. The descriptions of 36 G. arenarius syntypes indicate that males of this species have 55 pairs of legs, whereas females have 55 to 59 leg pairs, with 57 as the most common number.
Geophilus admarinus is a species of soil centipede in the family Geophilidae. It's found in southeast Alaska under stones near the low tide mark and is capable of surviving prolonged submersion underwater.
Geophilus alaskanus is a species of soil centipede in the family Geophilidae found in Alaska. It bears similarities to Mecistocephalus attenuatus, however unlike M. attenuatus, it has oblong cephalic lamina, unarmed coxae, and unarmed claw at base. G. alaskanus is dull orange-brown in color, grows to about 30 millimeters, and has 53 leg pairs with the first pair being very small and the last pair being distinctly larger than the others. It also possesses coalesced frontal lamina, unexposed basal lamina broader than the cephalic, a prehensorial sternum that's broader than it is long, distinct lateral grooves, obsolete prosternal teeth, a smooth, bare dorsal surface except for two faint impressed lines, and a small pleurae of the last segment marked with 8–10 pigmented pores of which the posterior one of the ventral face is larger than the others and sunk in a deep cavity.
Geophilus gracilis is a species of soil centipede in the family Geophilidae found most commonly in Britain and Ireland, though specimens have also been recorded in Chile, France, Greece, and Algeria. It lives under mud and stones along the coast near or below the high tide mark, grows up to 30 millimeters in length, and is bright yellow in color with a darker reddish head. Males of this species have 51 to 57 pairs of legs; females have 51 to 61 leg pairs. This species is often confused with G. flavus and, in coastal locations, G. osquidatum.
Geophilus mordax, also called the pitted soil centipede, is a species of soil centipede in the family Geophilidae found in North America, especially Indiana, Arkansas, Louisiana, North and South Carolina, and Florida. It grows up to 50 millimeters in length, though it averages 25–40, has 49–53 leg pairs in males and 49–57 in females, and is bright red in color. G. mordax also bears 3–5 labral teeth, a short and robust apical claw of the second maxillae, an exposed prebasal plate, and a large, consolidated sacculi.
Geophilus virginiensis is a species of soil centipede in the family Geophilidae found in Virginia. It grows up to 50 millimeters in length, has 49–57 leg pairs, and is tawny yellow in color.
Geophilus ampyx is a species of soil centipede in the family Geophilidae found in North America, especially South Carolina. It grows up to 52 millimeters in length, though it averages 30–40, has 49–53 leg pairs in males and 51–55 in females, and is bright red in color. G. ampyx also bears five sclerotized and deeply pigmented labral teeth and a nearly entirely exposed prebasal plate. It's often confused with G. mordax, though it can be differentiated by the absence of sacculi.
Geophilus cayugae is a species of soil centipede in the family Geophilidae found in New York, Virginia, and North Carolina. It grows up to 68 millimeters in length, though it averages 25–40, has 57–65 leg pairs in males and 63–67 in females, and is bright whitish yellow in color. G. cayugae can be identified by its well-developed paxilli, weakly consolidated sacculi, and concealed prebasal plate combined with lack of consolidated prebasal areas. It's found beneath rocks and other debris in high elevation montane regions.
Geophilus oweni is a species of soil centipede in the family Geophilidae found in Missouri, Indiana, and Ohio. It grows up to 40 millimeters in length, has 67-73 leg pairs in males and 71-77 in females, and varies from bright to faded yellow in color. G. oweni can be identified by its numerous leg pairs, lack of consolidated sacculi, prelabral consolidated areas, and long, thin second maxillary apical claws. G. missouriensis was found to be synonymous with G. oweni.
Geophilus vittatus, also known as the diamondback soil centipede, is a species of soil centipede in the family Geophilidae widespread in North America and named for the dark band of diamond-shaped markings across its back. G. vittatus grows up to 52 millimeters in length, though it averages 25–40, and ranges from brown- to orange-yellow in color, with 49–53 leg pairs in females, 49–51 leg pairs in males, and a lack of consolidated carpophagus structures. It can be found under any debris on the forest floor, but mostly under loose bark and occasionally near the sea.
Geophilus varians is a species of soil centipede in the family Geophilidae found in North America, particularly from South Carolina to Indiana, Ohio, Pennsylvania, and Virginia. It grows up to 40 millimeters, though it averages 30–35, ranges in color from light faded orange to yellow or whitish yellow with a deeper and brighter head, and has 53–59 leg pairs in males and 55–61 in females, as well as a complete lack of consolidated paxilli and sacculi, concealed prebasal plate, and unusually long ultimate legs.
Geophilus aztecus is a species of soil centipede in the family Geophilidae found in Mexico and Guatemala. The original description of this species is based on a dark red specimen measuring 44 mm in length with antennae moniliform at the apex. This species can reach 52 mm in length and has 57 or 59 pairs of legs.
Geophilus truncorum is a species of soil centipede in the family Geophilidae found across Western Europe, though it reaches as far as Poland, Italy, and Morocco. This centipede is relatively small, growing up to 20mm in length, with a yellow or orangeish brown body and dark yellow or brown head, denser and shorter hair than most Geophilus species, a main plate almost as elongated as in G. flavus (115:100), and distinct carpophagus fossae on the anterior sternites. Males of this species have 35 to 41 pairs of legs; females have 37 to 41.
Geophilus pusillus is a species of soil centipede in the family Geophilidae found in Algeria. It grows up to 11 millimeters in length. The original description of this species is based on ten specimens: five male specimens from Algeria with 31 pairs of legs, and five specimens from Germany. Records from the Alpstein mountains indicate that G. pusillus is a soil-dwelling species that prefers humus-rich soil, but these records deserve confirmation. Some authorities consider the specimens recorded from Europe to be specimens of G. ribauti that have been misidentified as specimens of the Algerian species G. pusillus.
Geophilus crenulatus is a species of soil centipede in the family Geophilidae found in a ravine above the Hemis monastery in India, 3525 meters above sea level. It was originally thought to be a subspecies of G. intermissus and named Geophilus intermissus var. crenulata. The original description of this species is based on a specimen measuring 22 mm in length with 57 pairs of legs.
