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Harpetida is one of the eleven orders of the extinct arthropod class Trilobita. The first harpetid trilobites appear in the Upper Cambrian, and the last species die out at the end of the Devonian period.
Phacopida ("lens-face") is an order of trilobites that lived from the Late Cambrian to the Late Devonian. It is made up of a morphologically diverse assemblage of taxa in three related suborders.
The Late Devonian extinction consisted of several extinction events in the Late Devonian Epoch, which collectively represent one of the five largest mass extinction events in the history of life on Earth. The term primarily refers to a major extinction, the Kellwasser event, also known as the Frasnian-Famennian extinction, which occurred around 372 million years ago, at the boundary between the Frasnian stage and the Famennian stage, the last stage in the Devonian Period. Overall, 19% of all families and 50% of all genera became extinct. A second mass extinction called the Hangenberg event, also known as the end-Devonian extinction, occurred 359 million years ago, bringing an end to the Famennian and Devonian, as the world transitioned into the Carboniferous Period.
The Frasnian is one of two faunal stages in the Late Devonian Period. It lasted from 382.7 million years ago to 372.2 million years ago. It was preceded by the Givetian Stage and followed by the Famennian Stage.
The Famennian is the later of two faunal stages in the Late Devonian Epoch. The most recent estimate for its duration estimates that it lasted from around 371.1 million years ago to 359.3 million years ago. An earlier 2012 estimate, still used by the International Commission on Stratigraphy, estimated that it lasted from 372.2 million years ago to 358.9 million years ago. It was preceded by the Frasnian stage and followed by the Tournaisian stage.
Chotecops is a genus of trilobites from the order Phacopida, suborder Phacopina, family Phacopidae. It was initially erected as a subgenus of Phacops but some later authors thought it distinctive enough to raise its status. Species assigned to this genus occur between the Emsian and the Famennian. Chotecops is the most abundant trilobite in the Hunsrück Slate and due to the excellent preservation, often soft tissue such as antennae and legs have been preserved as a thin sheet of pyrite.
Atrypa is a genus of brachiopod with round to short egg-shaped shells covered with many fine radial ridges. Growth lines form perpendicular to the costae and are spaced approximately 2 to 3 times further apart than the costae.. The pedunculate valve is slightly convex, but oftentimes levels out or becomes slightly concave toward the anterior margin. The brachial valve is highly convex. Neither valve contains an interarea. Atrypa had a large geographic range and occurred from the late Lower Silurian (Telychian) to the early Upper Devonian (Frasnian). Other sources expand the range from the Late Ordovician to Carboniferous, approximately from 449 to 336 Ma. A proposed new species, A. harrisi, was found in the trilobite-rich Floresta Formation in Boyacá, Colombia.
Leonaspis is a widespread genus of odontopleurid trilobite that lived from the Late Ordovician to the late Middle Devonian. Fossils of various species have been found on all continents except Antarctica.
Cyphaspis is a genus of small trilobite that lived from the Late Ordovician to the Late Devonian. Fossils have been found in marine strata in what is now Europe, Africa and North America. Various species had a compact body, and a large, bulbous glabellum. Many species had long spines arranged similarly to closely related genera, such as Otarian, Otarionella, Chamaeleoaspis, and Namuropyge.
Harpides is an extinct genus of harpetid trilobite of the family Harpididae.
Mcnamaraspis is an extinct monospecific genus of arthrodire placoderm that inhabited the ancient reef system of north Western Australia during the Frasnian epoch of the Late Devonian period. The type specimen was found and described by John A. Long from the Gogo Formation near Fitzroy Crossing. This fossil fish showed new anatomical features in arthrodires, like the well-preserved annular (ring-shaped) cartilages of the snout, previously inferred to be present by Erik Stensiö of Sweden. It is occasionally referred to as "The Gogo Fish" after the locale the holotype was excavated from.
Odontopleurida is an order of very spinose trilobites closely related to the trilobites of the order Lichida. Some experts group the Odontopleurid families, Odontopleuridae and Damesellidae, within Lichida. Odontopleurids tend to have convex, bar-shaped cephalons, and lobed, knob-shaped glabella that extend to, or almost to the anterior margin. Many, if not almost all odontopleurids have long spines that are derived either from the margins of the exoskeleton, or from granular or tubercular ornamentation, or both. Many odontopleurids are so spinose so as to be described as having "spines on (their) spines." Odontopleurids have 8 to 13 thoracic segments, with Odontopleuridae odontopleurids having no more than 10, and Damesellidae odontopleurids having no more than 13. The pygidium tends to be very small, and invariably has long spines emanating from it in all known genera.
Odontopleuridae is a family of odontopleurid trilobites found in marine strata throughout the world. Odontopleurids of Odontopleuridae first appear in Late Cambrian-aged marine strata, and the last genera perish by the end of the Frasnian stage during the Late Devonian. The members of Odontopleuridae are famous for their spinose appearance, having long, often numerous spines along the edges of their exoskeletons, and derived from ends of segments or tubercle ornaments.
Acuticryphops is a genus of trilobite that lived during the late Frasnian, particularly between the Lower and Upper Kellwasser horizons. It is known from mid-European Avalonia and Armorica and the North Gondwana margin.
Entomaspis is an extinct genus of harpetid trilobite from Upper Cambrian to Early Ordovician marine strata of the United States. Species are typified by their proportionally large, vaulted, croissant-shaped or bonnet-shaped cephalons that have the cheeks freed to become elongated, curved librigenial spines, and by their comparatively large, crescent-shaped eyes.
Librostoma is a subclass of trilobites defined by having a natant hypostome, which is a hypostome that is free from the anterior doublure and aligned with the anterior of the glabella, this is unlike a conterminant hypostome, which is attached to the exoskeleton.
Magreanops is a phacopid trilobite in the family Phacopidae. The type species is Magreanops renateae and the genus also includes Magreanops monachus, both described from the Frasnian-aged Bovesse Formation. and Moulin Liénaux Formations in Belgium during 2018.
Elginerpetontidae is an extinct family of basal stegocephalian tetrapodomorphs which lived in Europe during the Frasnian stage of the Late Devonian epoch. It contains the genera Elginerpeton, Obruchevichthys and Webererpeton.
Dudleyaspis is an extinct genus of Lower to Middle Devonian odontopleurid trilobites that lived in a shallow sea that lay between Euramerica and Gondwana. It was named in 1949 by Prantl & Pribyl.
Eskoharpes is a genus of harpetid trilobites. Along with Globoharpes it is one of the only harpetids that are found from the Frasnian stage of the Devonian period; it most likely evolved from Lioharpes.
References
- ↑ McNamara, Kenneth; Feist, Raimund; Ebach, Malte (2009). "Patterns of Evolution and Extinction in the Last Harpetid Trilobites During the Late Devonian (Frasnian)". Palaeontology. 52: 11–33. Bibcode:2009Palgy..52...11M. doi:10.1111/j.1475-4983.2008.00832.x. S2CID 129912220.
