Agnostida are an order of extinct arthropods which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician.
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.
The pygidium is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is composed of fused body segments, sometimes with a tail, and separated from thoracic segments by an articulation.
Phacopida ("lens-face") is an order of trilobites that lived from the Late Cambrian to the Late Devonian. It is made up of a morphologically diverse assemblage of taxa in three related suborders.
Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half of the Lower Cambrian, and the last species did not survive the Ordovician–Silurian extinction event.
Lichida is an order of typically spiny trilobite that lived from the Furongian to the Devonian period. These trilobites usually have 8–13 thoracic segments. Their exoskeletons often have a grainy texture or have wart or spine-like tubercles. Some species are extraordinarily spiny, having spiny thoracic segments that are as long or longer than the entire body, from cephalon (head) to pygidium (tail). The sections of the pygidia are leaf-like in shape and also typically end in spines.
Phacopidae is a family of phacopid trilobites that ranges from the Lower Ordovician to the Upper Devonian, with representatives in all paleocontinents.
Dalmanites is a genus of trilobite in the order Phacopida. They lived from the Late Ordovician to Middle Devonian.
Deiphon is a distinctive genus of Silurian phacopid trilobites of the family Cheiruridae found in Western and Central Europe, and in Central and Eastern United States. The type species, D. forbesi, from England, Bohemia, and Sweden, was discovered and described by the French paleontologist, Joachim Barrande in 1850.
Morocconites malladoides is an average size trilobite, which lived during the Devonian period, in what is now southern Morocco. This species is assumed to be a close relative of Acastoides. The most conspicuous feature is the very long upcurved frontal medial spine, a bit like an avocet bill. It is the only known species in this genus.
Bumastus is an extinct genus of corynexochid trilobites which existed from the Early Ordovician period to the Late Silurian period. They were relatively large trilobites, reaching a length of 6 in (15 cm). They were distinctive for their highly globular, smooth-surfaced exoskeleton. They possessed well-developed, large compound eyes and were believed to have dwelled in shallow-water sediments in life.
Ptychoparia is a genus of ptychopariid trilobites, and is the type genus of the family Ptychopariidae, and the order Ptychopariida.
Odontochile is a genus of trilobites in the order Phacopida, family Dalmanitidae.
Librostoma is a subclass of trilobites defined by having a natant hypostome, which is a hypostome that is free from the anterior doublure and aligned with the anterior of the glabella, this is unlike a conterminant hypostome, which is attached to the exoskeleton.
Hollardops is a genus of trilobite in the order Phacopida that lived during the Devonian. Their fossils are found in the upper Emsian of western Europe and in the lower Emsian to lowermost Eifelian of North Africa. The type species, Asteropyge mesocristata, was described from Algeria by Le Maître in 1952. The genus Hollardops was erected by Morzadec in 1997. In the same year, Lieberman & Kloc erected Modellops and Philipsmithiana but those genera are regarded as subjective synonyms of Hollardops. The 10-segmented thoracic condition of Hollardops is a rare feature among acastid trilobites that almost always have 11 thorax segments. Van Viersen & Kloc (2022) revisited Hollardops and described a number of new species from the Devonian of Morocco. They also regarded Pennarbedops Bignon & Crônier, 2013 as a synonym of Hollardops. Van Viersen & Kloc construed Hollardops as a scavenger or predator with well-developed eyes, that used its shovel-like cephalon to plough the top layers of the sediment in search of food. Hollardops had small pits horizontally along the fringe of the exoskeleton; these are believed to have housed setae that allowed the trilobite to closely monitor its surroundings.
Eskoharpes is a genus of harpetid trilobites. Along with Globoharpes it is one of the only harpetids that are found from the Frasnian stage of the Devonian period. It most likely evolved from Lioharpes.
Globoharpes is a genus of harpetid trilobite known from the late Frasnian of Western Australia.
Lioharpes is a genus of trilobites in the order Harpetida. The genus Eskoharpes possibly evolved from it. It is synonymous with Fritchapsis.
Harpetidae is a family of trilobites in the order Harpetida. They first appear in the Furongian epoch. The Taghanic event at the end of the Middle Devonian would impact them severely, with no genera from before surviving to the Frasnian, where two new genera, Eskoharpes and Globoharpes appear, before going extinct themselves in the Kellwasser event. Globoharpes may be a synonym of Eskoharpes.
