Calymene

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Calymene
Temporal range: Tremadocian-Pragian
~489–409  Ma
Calymene clavicula dorsal.jpg
Calymene clavicula 32mm, Henryhouse Formation, Oklahoma, Cayugan
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Phacopida
Family: Calymenidae
Genus: Calymene
Brongniart, 1822
Type species
Calymene blumenbachii
Brongniart, 1822  [1]

Calymene Brongniart, 1822, is a genus of trilobites in the order Phacopida, suborder Calymenina, that are found throughout North America, North Africa, and Europe in primarily Silurian outcrops. [2] Calymene is closely related to Flexicalymene , and both genera are frequently found enrolled. [3] Calymene trilobites are small, typically 2 cm in length. The cephalon is the widest part of the animal and the thorax usually has 13 segments. [4]

Contents

The correct genus authorship is Brongniart (1822). [5] A previously published genus description in Desmarest (1816) [6] (often mis-cited as "Calymena" Desmarest, 1817) was suppressed by ICZN Opinion 1433.

Etymology

Calymene - meaning beautiful crescent as a reference to the glabella.

Known species and locations

Reassigned species

Since the genus Calymene was established early on in paleontology, a number of species previously assigned to it have since been transferred to other genera: [9]

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<span class="mw-page-title-main">Silurian</span> Third period of the Paleozoic Era, 443–419 million years ago

The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.

<span class="mw-page-title-main">Trilobite</span> Class of extinct, Paleozoic arthropods

Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.

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<i>Phacops</i> Genus of arthropods (fossil)

Phacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian, with a broader time range described from the Late Ordovician. It was a rounded animal, with a globose head and large eyes, and probably fed on detritus. Phacops is often found rolled up ("volvation"), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus.

<i>Flexicalymene</i> Genus of trilobites (fossil)

Flexicalymene Shirley, 1936. is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and Family Calymenidae. Flexicalymene specimens can be mistaken for Calymene, Gravicalymene, Diacalymene and a few other Calymenina genera. They are used as an index fossil in the Ordovician. Ohio and North America are particularly known for being rich with Flexicalymene fossils.

<i>Gravicalymene</i> Extinct genus of trilobites

Gravicalymene Shirley, 1936, is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and family Calymenidae. Species included in this genus have previously been allocated to Calymene Brongniart 1822,Flexicalymene Shirley, 1936. and Sthenarocalymene Siveter 1977.

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Asaphus is a genus of trilobites that is known from the Lower and Middle Ordovician of northwestern Europe.

<i>Calymene blumenbachii</i> Extinct species of trilobite

Calymene Brongniart in Desmarest (1817), sometimes erroneously spelled blumenbachi, is a species of trilobite discovered in the limestone quarries of the Wren's Nest in Dudley, England. Nicknamed the Dudley Bug or Dudley Locust by 18th-century quarrymen it became a symbol of the town and featured on the Dudley County Borough Council coat-of-arms. Calymene blumenbachii is commonly found in Silurian rocks and is thought to have lived in the shallow waters of the Silurian, in low-energy reefs. This particular species of Calymene is unique to the Wenlock series in England, and comes from the Wenlock Limestone Formation in Much Wenlock and the Wren's Nest in Dudley. These sites seem to yield trilobites more readily than any other areas on the Wenlock Edge, and the rock here is dark grey as opposed to yellowish or whitish as it appears on other parts of the Edge, just a few miles away, in Church Stretton and elsewhere. This suggests local changes in the environment in which the rock was deposited.

<i>Balizoma</i> Genus of trilobites

Balizoma is a genus of trilobites from the family Encrinuridae established by David J. Holloway in 1980. It has only been found in rocks of Silurian age. Its type species, B. variolaris, is currently the only named species of the genus, and is found in England. The neotype of B. variolaris was collected from the Much Wenlock Limestone Formation at Dudley, West Midlands. That specimen was first illustrated in Sir Roderick Impey Murchison's classic book, The Silurian System. B. variolaris was the original "strawberry-headed" trilobite of Dudley, so-named because of its nodular glabellar tubercles, and well known to early trilobite collectors. Additional species were originally assigned to Balizoma, but were subsequently placed in other encrinurine genera.

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References

  1. 1 2 Derek J. Siveter (1985). "The type species of Calymene (Trilobita) from the Silurian of Dudley, England" (PDF). Palaeontology . 28 (4): 783–792. Archived from the original (PDF) on 2017-10-16. Retrieved 2017-10-16.
  2. "†Calymene Muenster 1840 (trilobite)". The Paleontology Database. Archived from the original on 2012-07-20. Retrieved April 29, 2012.
  3. John P. Rafferty, ed. (2010). The Paleozoic Era: Diversification of Plant and Animal Life. Geologic History of Earth. Britannica Educational Publishing. ISBN   978-1-61530-196-6.
  4. Milson, C.; Rigby, S. (2004). Fossils at a Glance. Oxford: Blackwell Science Ltd.
  5. Brongniart, A. 1822. Les Trilobites. pp. 1-65, pls. 1-4 in: Histoire Naturelle des Crustacés Fossiles. Paris. F.-G. Levrault, Libraire. 154 pp. 11 pls.
  6. Desmarest, A-G. 1816. Calymène. pp. 49-50 in: Nouveau Dictionnaire d'Histoire Naturelle, Nouvelle Edition, Tome 5.
  7. "Calymene celebra Raymond 1916 (trilobite)". Paleobiology Database. Retrieved 20 August 2011.
  8. Alex J. Chestnut. "Using morphometrics, phylogenetic systematics and parsimony analysis to gain insight into the evolutionary affinities of the Calymenidae Trilobita". OhioLINK ETD Center. Retrieved August 21, 2011.
  9. Moore, R. C. (1959). Arthropoda I - Arthropoda General Features, Proarthropoda, Euarthropoda General Features, Trilobitomorpha. Treatise on Invertebrate Paleontology. Vol. Part O. Boulder, Colorado/Lawrence, Kansas: Geological Society of America/University of Kansas Press. pp. 1–560. ISBN   0-8137-3015-5.
  10. 1 2 Esteve, Jorge (2015). "Systematic revision of the genus Solenopleura Angelin, 1854, Ptychopariida, Trilobita, Cambrian Series 3". Annales de Paléontologie. 101 (3): 185–192. Bibcode:2015AnPal.101..185E. doi:10.1016/j.annpal.2015.05.001.
  11. Pärnaste, Helje (2004). "Revision of the Ordovician cheirurid trilobite genus Reraspis with the description of the earliest representative" (PDF). Proc. Estonian Acad. Sci. Geol. 53 (2): 125–138. doi:10.3176/geol.2004.2.03. S2CID   55808470.
  12. Ramskjöld, L.; Werdelin, L. (1991). "The phylogeny and evolution of some phacopid trilobites". Cladistics. 7: 29–74. doi: 10.1111/j.1096-0031.1991.tb00021.x .