Calymene | |
---|---|
Calymene clavicula 32mm, Henryhouse Formation, Oklahoma, Cayugan | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | † Trilobita |
Order: | † Phacopida |
Family: | † Calymenidae |
Genus: | † Calymene Brongniart, 1822 |
Type species | |
Calymene blumenbachii Brongniart, 1822 [1] |
Calymene Brongniart, 1822, is a genus of trilobites in the order Phacopida, suborder Calymenina, that are found throughout North America, North Africa, and Europe in primarily Silurian outcrops. [2] Calymene is closely related to Flexicalymene , and both genera are frequently found enrolled. [3] Calymene trilobites are small, typically 2 cm in length. The cephalon is the widest part of the animal and the thorax usually has 13 segments. [4]
The correct genus authorship is Brongniart (1822). [5] A previously published genus description in Desmarest (1816) [6] (often mis-cited as "Calymena" Desmarest, 1817) was suppressed by ICZN Opinion 1433.
Calymene - meaning beautiful crescent as a reference to the glabella.
Since the genus Calymene was established early on in paleontology, a number of species previously assigned to it have since been transferred to other genera: [9]
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
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Phacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian, with a broader time range described from the Late Ordovician. It was a rounded animal, with a globose head and large eyes, and probably fed on detritus. Phacops is often found rolled up ("volvation"), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus.
Flexicalymene Shirley, 1936. is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and Family Calymenidae. Flexicalymene specimens can be mistaken for Calymene, Gravicalymene, Diacalymene and a few other Calymenina genera. They are used as an index fossil in the Ordovician. Ohio and North America are particularly known for being rich with Flexicalymene fossils.
Gravicalymene Shirley, 1936, is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and family Calymenidae. Species included in this genus have previously been allocated to Calymene Brongniart 1822,Flexicalymene Shirley, 1936. and Sthenarocalymene Siveter 1977.
Asaphus is a genus of trilobites that is known from the Lower and Middle Ordovician of northwestern Europe.
Calymene Brongniart in Desmarest (1817), sometimes erroneously spelled blumenbachi, is a species of trilobite discovered in the limestone quarries of the Wren's Nest in Dudley, England. Nicknamed the Dudley Bug or Dudley Locust by 18th-century quarrymen it became a symbol of the town and featured on the Dudley County Borough Council coat-of-arms. Calymene blumenbachii is commonly found in Silurian rocks and is thought to have lived in the shallow waters of the Silurian, in low-energy reefs. This particular species of Calymene is unique to the Wenlock series in England, and comes from the Wenlock Limestone Formation in Much Wenlock and the Wren's Nest in Dudley. These sites seem to yield trilobites more readily than any other areas on the Wenlock Edge, and the rock here is dark grey as opposed to yellowish or whitish as it appears on other parts of the Edge, just a few miles away, in Church Stretton and elsewhere. This suggests local changes in the environment in which the rock was deposited.
Balizoma is a genus of trilobites from the family Encrinuridae established by David J. Holloway in 1980. It has only been found in rocks of Silurian age. Its type species, B. variolaris, is currently the only named species of the genus, and is found in England. The neotype of B. variolaris was collected from the Much Wenlock Limestone Formation at Dudley, West Midlands. That specimen was first illustrated in Sir Roderick Impey Murchison's classic book, The Silurian System. B. variolaris was the original "strawberry-headed" trilobite of Dudley, so-named because of its nodular glabellar tubercles, and well known to early trilobite collectors. Additional species were originally assigned to Balizoma, but were subsequently placed in other encrinurine genera.
Bumastus is an extinct genus of corynexochid trilobites which existed from the Early Ordovician period to the Late Silurian period. They were relatively large trilobites, reaching a length of 6 in (15 cm). They were distinctive for their highly globular, smooth-surfaced exoskeleton. They possessed well-developed, large compound eyes and were believed to have dwelled in shallow-water sediments in life.
Paleontology in Oklahoma refers to paleontological research occurring within or conducted by people from the U.S. state of Oklahoma. Oklahoma has a rich fossil record spanning all three eras of the Phanerozoic Eon. Oklahoma is the best source of Pennsylvanian fossils in the United States due to having an exceptionally complete geologic record of the epoch. From the Cambrian to the Devonian, all of Oklahoma was covered by a sea that would come to be home to creatures like brachiopods, bryozoans, graptolites and trilobites. During the Carboniferous, an expanse of coastal deltaic swamps formed in areas of the state where early tetrapods would leave behind footprints that would later fossilize. The sea withdrew altogether during the Permian period. Oklahoma was home a variety of insects as well as early amphibians and reptiles. Oklahoma stayed dry for most of the Mesozoic. During the Late Triassic, carnivorous dinosaurs left behind footprints that would later fossilize. During the Cretaceous, however, the state was mostly covered by the Western Interior Seaway, which was home to huge ammonites and other marine invertebrates. During the Cenozoic, Oklahoma became home to creatures like bison, camels, creodonts, and horses. During the Ice Age, the state was home to mammoths and mastodons. Local Native Americans are known to have used fossils for medicinal purposes. The Jurassic dinosaur Saurophaganax maximus is the Oklahoma state fossil.
Bailiaspis Resser, 1936, is a Middle Cambrian (Miaolingian) trilobite genus belonging to the Family Conocoryphidae Angelin, 1854. Within the Acado-Baltic region, the genus ranges from Wuliuan into Guzhangian age strata.
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