Redlichiida

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Redlichiida
Temporal range: Cambrian Stage 3 – Middle Cambrian, 521–500  Ma
Te Li Si Duo Han Wu Zhi Hui Chong Hua Shi .jpg
Cambropallas telesto, from Morocco
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Redlichiida
Richter, 1932
Suborders

Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis (suborder Olenellina), and Lemdadella (suborder Redlichiina), both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.

Contents

Description

Olenellus chiefensis, suborder Olenellina, showing the visual surface has broken away, the lack of dorsal sutures, and the enlarged pleurae of the 3rd thorax segment from the front Olenellus chiefensis CRF.jpeg
Olenellus chiefensis, suborder Olenellina, showing the visual surface has broken away, the lack of dorsal sutures, and the enlarged pleurae of the 3rd thorax segment from the front

Most redlichiids are rather flat (or have low dorso–ventral convexity) and their exoskeleton typically has an oval outline, about 1½× longer than wide. Each back edge of the headshield (or cephalon) very often carries a spine, termed a genal spine. The eye lobes are sickle-shaped, long and extend from the frontal lobe of the central raised area of the cephalon (or glabella) curving outward and increasingly backwards and sometimes eventually inwards again. The visual surface, that contains the calcite lenses is surrounded by fracture lines (or circumocular sutures), so that it has most often broken away from the rest of the cephalon. The glabella tapers forward and is relatively long, with the frontal lobe boss-like or pointy, followed by three rings or pairs of lobes (defined by furrows that may or may not cross over the midline), and finally at the back of the cephalon the so-called occipital ring. On the ventral side of the cephalon, the palate (or hypostome) is attached to the part of the calcified exoskeleton that defines the contour at the ventral side (the so-called doublure), a state called conterminant. The hypostomes of redlichiids have narrow borders, are not split into backward pointing forks, and have only small muscle attachment areas (or maculae). The articulate middle part of the exoskeleton (or thorax) is composed of many of segments that often end in a spine at the side of the animal. To each side of the central axis, the third segment from the front may have an exta large and wide rib (a state called macropleural). The tailshield (or pygidium) is small but appears to be composed of more than one segment. In Olenellina the earliest larval stage (called protaspis) has not been found, so it is presumed it was not calcified. The development of Redlichia from protaspis to adult was gradual without a metamorphosis at any stage. [1]

Two major Lagerstätten where redlichiids are found are the Emu Bay shales of Southern Australia and the Maotianshan shales near Chengjiang in China. Other regions include Morocco, Labrador, Canada, the western United States, and Bohemia, Czech Republic.

The appendages have been preserved in a few specimens. They follow typical trilobite patterns in terms of the number, placement, and types of legs, antennae, gills, etc.

Suborders

The order Redlichiida is divided into two suborders: Olenellina and Redlichiina. The main difference between the two groups is the lack of facial sutures in the Olenellina. This absence of the facial sutures is regarded as primitive by most scholars. Opisthoparian facial sutures are a shared character of all Redlichiina. In other trilobites, dorsal sutures may be opisthoparian, gonatoparian, proparian or they may be lost secondarily. The absence in the fossil record of the earliest larval stage, the protaspid, suggests that it may have been uncalcified, which would be a second unique character that distinguishes the Olenellina from all other trilobites. [1]

Fossils of olenellinid trilobites are found in North America, and other associated areas that comprised the Cambrian continent of Laurentia. They are very common and are used to define the extent of Laurentia. Their abrupt disappearance marks the boundary between the Lower and the Middle Cambrian.

Fossils of redlichiinid are associated with Cambrian regions other than Laurentia.

Related Research Articles

<span class="mw-page-title-main">Trilobite</span> Class of extinct, Paleozoic arthropods

Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.

<i>Paradoxides</i> Extinct genus of trilobites

Paradoxides is a genus of large to very large trilobite found throughout the world during the Middle Cambrian period. One record-breaking specimen of Paradoxides davidis, described by John William Salter in 1863, is 37 cm (15 in). The cephalon was semicircular with free cheeks ending in long, narrow, recurved spines. Eyes were crescent shaped providing an almost 360° view, but only in the horizontal plane. Its elongate thorax was composed of 19–21 segments and adorned with longish, recurved pleural spines. Its pygidium was comparatively small. Paradoxides is a characteristic Middle-Cambrian trilobite of the 'Atlantic' (Avalonian) fauna. Avalonian rocks were deposited near a small continent called Avalonia in the Paleozoic Iapetus Ocean. Avalonian beds are now in a narrow strip along the East Coast of North America, and in Europe.

<span class="mw-page-title-main">Emuellidae</span> Extinct family of trilobites

Emuellidae are a small family of trilobites, a group of extinct marine arthropods, that lived during the late Lower Cambrian of the East Gondwana supercontinent, in what are today South-Australia and Antarctica.

<span class="mw-page-title-main">Olenellina</span> Extinct suborder of trilobites

Olenellina is a suborder of the order Redlichiida of trilobites that occurs about halfway during the Lower Cambrian, at the start of the stage called the Atdabanian. The earliest trilobites in the fossil record are arguably Olenellina, although the earliest Redlichiina,Ptychopariida, and Eodiscina follow quickly. The suborder died out when the Lower Cambrian passed into the Middle Cambrian, at the end of the stage called Toyonian. A feature uniting the Olenellina is the lack of rupture lines in the headshield, which in other trilobites assist the periodic moulting, associated with arthropod growth. Some derived trilobites have lost facial sutures again, but all of these are blind, while all Olenellina have eyes.

<span class="mw-page-title-main">Redlichiina</span> Extinct suborder of trilobites

Redlichiina is a suborder of the order Redlichiida of Trilobites. The suborder contains three superfamilies: Emuelloidea, Redlichioidea and Paradoxidoidea. These trilobites are some of the oldest trilobites known. They originated at the beginning of the Cambrian Period and disappeared at the end of the middle Cambrian.

<i>Olenelloides</i>

Olenelloides armatus is an extinct, small sized olenelloid redlichiid trilobite arthropod. It lived during the later part of the Botomian stage, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the Cambrian Period. The most conspicuous feature is the hexagonal head shield that carries 6 ray-like spines..

<i>Nephrolenellus</i>

Nephrolenellus is an extinct genus of trilobite, fossil marine arthropods, of relatively small size. Currently two species are attributed to it. Nephrolenellus lived at the end of the Lower Cambrian. Species are known from the Great Basin of California, Nevada and Arizona, with one specimen from Canada.

<i>Mesonacis</i>

Mesonacis is an extinct genus of trilobite that lived during the Botomian, found in North-America, and the United Kingdom. Some of the species now regarded part of Mesonacis, have previously been assigned to Angustolenellus or Olenellus (Angustolenellus). Angustolenellus is now regarded a junior synonym of Mesonacis.

<i>Fallotaspis</i> Cambrian Period genus of trilobites

Fallotaspis is a genus of redlichiid trilobites found in Early Cambrian-aged strata of the United States and Morocco.

The cephalon is the head section of an arthropod. It is a tagma, i.e., a specialized grouping of arthropod segments. The word cephalon derives from the Greek κεφαλή (kephalē), meaning "head".

<span class="mw-page-title-main">Holmiidae</span> Family of trilobites

Holmiidae is a family of trilobites, that lived during the Lower Cambrian (Atdabanian). The Holmiidae is a diverse family of eight genera containing at least 17 species. It includes some of the earliest trilobites of Baltica. Holmiidae occur throughout Baltica and Western Laurentia, and also in Morocco.

<i>Biceratops</i>

Biceratops is an extinct genus of olenelloid redlichiid trilobites, of average size, with the largest specimen 8 centimetres or 3.1 inches long, not including the huge pleural spines of the 3rd segment of the thorax. It lived during the Toyonian stage, in what is today the South-Western United States. Biceratops can easily be distinguished from other members of Biceratopsidae by the absence of genal spines, in combination with effaced features of the raised axial area of the head shield, that is bordering the two horn-like projections that carry the eyes. Biceratops nevadensis is the only known species in this genus.

<span class="mw-page-title-main">Olenelloidea</span> Extinct superfamily of trilobites

The Olenelloidea are a superfamily of trilobites, a group of extinct marine arthropods. They lived during the late Lower Cambrian and species occurred on all paleocontinents.

Mesolenellus is an extinct genus of trilobites that lived during the lower Cambrian (Botomian), found in Greenland and Spitsbergen.

<i>Odontochile</i> Genus of trilobites

Odontochile is a genus of trilobites in the order Phacopida, family Dalmanitidae.

<i>Bristolia</i>

Bristolia is an extinct genus of trilobite, fossil marine arthropods, with eight or more small to average size species. It is common in and limited to the Lower Cambrian shelf deposits across the southwestern US, which constitutes part of the former paleocontinent of Laurentia.

<i>Emigrantia</i>

Emigrantia is an extinct genus of trilobites, fossil marine arthropods, of small to average size. It lived during the Toyonian stage, in what is today the South-Western United States. Emigrantia can easily be distinguished from other trilobites by the sturdy but not inflated genal spines, that are attached at midlength of the cephalon, in combination with effaced features of the raised axial area of the head shield.

Bolbolenellus is an extinct genus of trilobites, fossil marine arthropods, with five species attributed to it currently. It can be easily distinguished from all other trilobites by the combination of the absence of dorsal sutures in the head shield like all Olenellina, and a distinctly bulbous frontal lobe (L4) of the raised axial area in the head called glabella. The species lived at the end of the Lower Cambrian.

<span class="mw-page-title-main">Richterops</span>

Richterops is an extinct genus of trilobite arthropods. The genus lived during the middle of the Atdabanian or the early part of the Botomian stage, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the second half of the Lower Cambrian. It has been found in southern Morocco. It can be recognised by the long spines of the headshield that are a smooth continuation of the frontal edge, and the enlarged spines on the 11th segment of the thorax.

<span class="mw-page-title-main">Fallotaspidoidea</span>

The ”Fallotaspidoidea” are a superfamily of trilobites, a group of extinct marine arthropods. It lived during the Lower Cambrian (Atdabanian) and species occurred on all paleocontinents except for the Gondwana heartland. A member of this group, Profallotaspis jakutensis, has long been the earliest known trilobite, but recently the redlichiid Lemdadella has been claimed as occurring even earlier.

References

  1. 1 2 Whittington, H.B.; et al. (1997). Introduction, Order Agnostida, Order Redlichiida. Treatise on Invertebrate Paleontology. Vol. Part O.Revised, Volume 1 – Trilobita. Boulder, Colorado/Lawrence, Kansas: Geological Society of America/University of Kansas Press. pp.  400–481. ISBN   0-8137-3108-9.