Redlichiina

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Redlichiina
Temporal range: late Lower - Middle Cambrian
RedlichiidaTakooensis.png
Redlichia takooensis  , a member of the trilobite suborder Redlichiina.
Lower Cambrian Emu Shale
Kangaroo Island, South Australia
(© Dave Simpson)
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Redlichiida
Suborder: Redlichiina
Richter, 1932
Superfamilies

Redlichiina is a suborder of the order Redlichiida of Trilobites. The suborder contains three superfamilies: Emuelloidea, Redlichioidea and Paradoxidoidea. These trilobites are some of the oldest trilobites known. They originated at the beginning of the Cambrian Period and disappeared (possibly by evolving into members of the Ptychopariida order) at the end of the middle Cambrian.

Physiology

Diagram showing the cephalon, thorax and pygidium of an example trilobite. SamGonIII cepthopyg.png
Diagram showing the cephalon, thorax and pygidium of an example trilobite.

Cephalon: Opisthoparian facial sutures. Early forms tend to have tapering, conical glabella with furrows extending far backwards while later forms with glabella expanding forwards to inflated frontal lobe. The hypostome is conterminant (e.g., Redlichia) or natant (e.g., Dolerolenus). The rostral plate is narrower than in suborder Olenellina, bound by rostral and connective sutures.

Thorax: Fulcrate or non-fulcrate, typically with many segments (60+ in an Emmuelid). The axis is infrequently spine-bearing.

Pygidium: Typically small, but can be larger and with many segments.

Related Research Articles

Trilobite Class of extinct, Paleozoic arthropods

Trilobites are a group of extinct marine artiopodan arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period, and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 300 million years.

Redlichiida

Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.

Ptychopariida

Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half of the Lower Cambrian, and the last species did not survive the Ordovician–Silurian extinction event.

Phacopina

The Phacopina comprise a suborder of the trilobite order Phacopida. Species belonging to the Phacopina lived from the Lower Ordovician (Tremadocian) through the end of the Upper Devonian (Famennian). The one unique feature that distinguishes Phacopina from all other trilobites are the very large, separately set lenses without a common cornea of the compound eye.

Emuellidae

Emuellidae are a small family of trilobites, a group of extinct marine arthropods, that lived during the late Lower Cambrian of the East Gondwana supercontinent, in what are today South-Australia and Antarctica. Emuellidae can be recognized among trilobites in having a set of unique features. The headshield or cephalon has large genal spines reaching back as far as the 3rd to 6th segment of the thorax. The eye-ridges contact the back of the frontal lobe of the glabella and extend laterally and backwards, roughly parallel to the frontal and lateral rim of the cephalon. There are small, clearly incised pits at the junction between the eye-ridge and the frontal lobe of the cephalic axis. The thorax reaches its greatest width at the 6th segment. The frontal part or prothorax consists of 6 segments, with number 5 and 6 fused, and the 6th carrying very large trailing spines. The rear part or opistothorax consists of a variable but extremely large number of segments.

<i>Emuella</i>

Emuella is a genus of trilobites of the family Emuellidae. Its fossils have been found in South Australia. It can be recognised by touching glabella and frontal border, and the sub-pentagonal head, as compared to, a short field between the front of the axis in the head or glabella and the border ridge, and a semi-circular headsheald in the sister-genus Balcoracania. Both emuellid genera share eye ridges that are positioned parallel to the frontal and lateral border of the head, prominent genal spines that are a smooth continuation of the lateral margin of the head, a prothorax of 6 segments, with the 5th and 6th merged and carrying large trailing spines. Both genera have in adulthood a highly variable but large number of segments of the opistothorax, although the largest number found in B. dailyi with 97 is much larger than in Emuella (52).

<i>Redlichia takooensis</i>

Redlichia takooensis is a species of redlichiid trilobite from the lower Cambrian-aged Emu Shale of Kangaroo Island, Australia.

<i>Estaingia bilobata</i>

Estaingia bilobata is a species of trilobite from the lower Cambrian period. Their fossils are found chiefly in Australia.

Olenellina

Olenellina is a suborder of the order Redlichiida of trilobites that occurs about halfway during the Lower Cambrian, at the start of the stage called the Atdabanian. The earliest trilobites in the fossil record are arguably Olenellina, although the earliest Redlichiina and Eodiscina follow quickly. The suborder died out when the Lower passed into the Middle Cambrian, at the end of the stage called Toyonian. A feature uniting the Olenellina is the lack of rupture lines in the headshield, which in other trilobites assist the periodic moulting, associated with arthropod growth. Some derived trilobites have lost facial sutures again, but all of these are blind, while all Olenellina have eyes.

<i>Kootenia</i>

Kootenia is a genus of trilobites of the family Dorypygidae. 118 specimens of Kootenia are known from the Greater Phyllopod bed, where they comprise 0.22% of the community. Its major characteristics are that of the closely related Olenoides, including medium size, a large glabella, and a medium-sized pygidium, but also a lack of the strong interpleural furrows on the pygidium that Olenoides has.

Redlichia

Redlichia is a genus of redlichiid trilobite in the family Redlichiidae, with large to very large species. Fossils of various species are found in Lower Cambrian (Toyonian)-aged marine strata from China, Korea, Pakistan, the Himalayas, Iran, Spain, southern Siberia, and Antarctica, and from Middle Cambrian (Ordian)-aged marine strata of Australia.

Emu Bay Shale

The Emu Bay Shale is a geological formation in Emu Bay, South Australia, containing a major Konservat-Lagerstätte. It is one of two in the world containing Redlichiidan trilobites. The Emu Bay Shale is dated as Cambrian Series 2, Stage 4, correlated with the upper Botomian Stage of the Lower Cambrian.

<i>Acidiscus</i>

Acidiscus Rasetti, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida Salter (1864). It lived during the Botomian stage = late Lower Cambrian Stage 4 ; the upper Botomian boundary corresponds to base of the Middle Cambrian, Miaolingian Series and Wuliuan stage.

<i>Holyoakia</i>

Holyoakia is a genus of very small trilobites of the family Dorypygidae, from the late Lower Cambrian of South Australia and Antarctica.

<i>Yunnanocephalus</i>

Yunnanocephalus is a genus of ptychopariid trilobite. It lived during the late Atdabanian and Botomian stages, in what are currently Antarctica, Australia and China. It was a "moderately common" member of the Chengjiang Fauna. Yunnanocephalus is the only genus currently assigned to the Yunnanocephalidae family.

<i>Tsunyidiscus</i>

Tsunyidiscus is a trilobite belonging to the Suborder Eodiscina. Tsunyidiscus appeared near the end of the Lower Cambrian, during the late Atdabanian stage of geologic time and some collections suggest it may have survived into the Botomian. The genus is very small, occulatre and isopypous with a narrow dome-shaped glabella and a narrow bullet-shaped pygidial axis. Thorax consists of three segments. Tsunyidiscus is the only genus currently attributed to the family Tsunyidiscidae.

<i>Nephrolenellus</i>

Nephrolenellus is an extinct genus of trilobite, fossil marine arthropods, of relatively small size. Currently two species are attributed to it. Nephrolenellus lived at the end of the Lower Cambrian. Species are known from the Great Basin of California, Nevada and Arizona, with one specimen from Canada.

<i>Pagetia</i>

Pagetia Walcott, 1916. is a small genus of trilobite, assigned to the Eodiscinid family Pagetiidae and which had global distribution during the Middle Cambrian. The genus contains 55 currently recognized species, each with limited spatial and temporal ranges.

The cephalon is the head section of an arthropod. It is a tagma, i.e., a specialized grouping of arthropod segments. The word cephalon derives from the Greek κεφαλή (kephalē), meaning "head".

Eodiscina

Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.

References