Agnostida are an order of extinct arthropods which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician.
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.
Nektaspida is an extinct order of non-mineralised artiopodan arthropods. They are known from the mid-Cambrian to the upper Silurian. Originally classified as trilobites, which they superficially resemble, they are now placed as close relatives as members of the Trilobitomorpha within Artiopoda. The order is divided into three major families; Emucarididae, Liwiidae, and Naraoiidae.
Harpetida is one of the eleven orders of the extinct arthropod class Trilobita. The first harpetid trilobites appear in the Upper Cambrian, and the last species die out at the end of the Devonian period.
Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies, but no suborders. Asaphids comprise some 20% of described fossil trilobites.
Phacopida ("lens-face") is an order of trilobites that lived from the Late Cambrian to the Late Devonian. It is made up of a morphologically diverse assemblage of taxa in three related suborders.
Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half of the Lower Cambrian, and the last species did not survive the Ordovician–Silurian extinction event.
Proetida is an order of trilobite that lived from the Ordovician to the Permian. It was the last surviving order of trilobite, dying out in the Permian-Triassic extinction event.
The Phacopina comprise a suborder of the trilobite order Phacopida. Species belonging to the Phacopina lived from the Lower Ordovician (Tremadocian) through the end of the Upper Devonian (Famennian). The one unique feature that distinguishes Phacopina from all other trilobites are the very large, separately set lenses without a common cornea of the compound eye.
Olenellina is a suborder of the order Redlichiida of trilobites that occurs about halfway during the Lower Cambrian, at the start of the stage called the Atdabanian. Olenellina are arguably the earliest trilobites in the fossil record as members of Redlichiina, although Ptychopariida and Eodiscina follow soon after. The suborder died out when the Lower Cambrian passed into the Middle Cambrian, at the end of the stage called Toyonian. A feature uniting the Olenellina is the lack of rupture lines in the headshield, which in other trilobites assist the periodic moulting, associated with arthropod growth. Some derived trilobites have lost facial sutures again, but all of these are blind, while all Olenellina have eyes.
Walliserops is a genus of spinose phacopid trilobite, of the family Acastidae, found in Lower to Middle Devonian age rocks from the Anti-Atlas Mountains of Morocco. All species of Walliserops possess a three-pronged "trident" that protrudes from the glabella. Walliserops is most closely related to the genus Comura.
Trimerocephalus is a genus of eyeless trilobites from the order Phacopida, family Phacopidae. It lived during the final stage of the Devonian, the Famennian, and became extinct at the end of this stage, together with all other trilobites with the exception of some Proetida. It can be found in Australia, the Czech Republic, France, Germany, Iran, Kazakhstan, Morocco, Poland, the Russian Federation (Urals), Spain, and the United Kingdom (England).
Aulacopleura is a genus of proetid trilobite that lived from the Middle Ordovician to the Middle Devonian. Some authors may classify this group as subgenus Otarion (Aulacopleura). The cephalon is semicircular or semielliptical, with border and preglabellar field. The glabella is short, with or without defined eye ridges connecting it with eyes of variable size. Spines at the rear outer corners of the cephalon are present, typically reaching back to the 2nd to 4th thorax segment. The 'palate' is not connected to the dorsal shield of the cephalon. The cephalon is pitted, or has small tubercles. The thorax has up to 22 segments. The pleural ends are usually rounded. The pygidium is small (micropygous), with an even margin. A. koninckii had a modern type of compound eye.
The hypostome is the hard mouthpart of trilobites found on the ventral side of the cephalon (head). The hypostome can be classified into three types based on whether they are permanently attached to the rostrum or not and whether they are aligned to the anterior dorsal tip of the glabella.
Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.
Cyclopygidae is a family of asaphid trilobites from the Ordovician. Cyclopygids had an extratropical distribution, and there is evidence that they lived in darker parts of the water column. Cyclopygids are characterized by enlarged eyes, with a wide angle of view, both horizontal and vertical, reminiscent of the eyes of dragonflies. These typically touch the glabella directly on the side. Cyclopygids all lack genal spines, but Symphysops carries a forward directed frontal spine on the glabella. It is presumed that at least the members of the genus Pricyclopyge swam upside down and had bioluminescent organs on the third thorax segment. Cyclopygids had between 7 and 5 thorax segments, a wide and stout axis, and short side lobes.
Typhlokorynetes plana is a species of small, button-shaped asaphid trilobites of the family Raphiophoridae that lived during the Early Tremadocian of Vermont, United States.
Trinucleioidea is a superfamily of trilobites. Traditionally placed within the Asaphida, it is now sometimes considered its own order, Trinucleida.
Lioharpes is a genus of trilobites in the order Harpetida. The genus Eskoharpes possibly evolved from it. It is synonymous with Fritchapsis.
