Ampyx (trilobite)

Last updated

Ampyx
Temporal range: Ordovician-Silurian
~478.6–416.0  Ma
Ampyx cetsarum - specimen number MGL 96718 - white (cropped).jpg
Ampyx cetsarum group
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Asaphida
Family: Raphiophoridae
Genus: Ampyx
Dalman, 1827
Type species
Ampyx nasutus
Species
  • A. gongwusuensisLee et al., 2016
  • A. nasutusDalman, 1827
Synonyms

Brachyampyx

Ampyx is an Ordovician-Silurian genus of Asaphid trilobites of the family Raphiophoridae. Species of Ampyx are characterized by three extended spines on the head-shield, one spine derived from each free cheek, and one spine emanating from the glabellum. [1] Species include Ampyx linleyensis (Lanvirn-Caradoc series). [2]

Contents

Species of Ampyx grew to an average length of 1.5 inches (3.8 cm).

Collective behaviour

Linear cluster of
Ampyx priscus Linear clusters of Ampyx priscus 2.png
Linear cluster of
Ampyx priscus

Fossils of the trilobite Ampyx priscus, dating back about 480 million years ago, have been recently described as clustered in lines along the ocean floor. The animals were all mature adults, and were all facing the same direction as though they had formed a conga line or a peloton. It has been suggested they line up in this manner to migrate, much as spiny lobsters migrate in single-file queues. Or perhaps they are getting together for mating. The findings suggest animal collective behaviour has very early evolutionary origins. [3] [4]

Distribution

Fossils of Ampyx have been found in: [5]

Ordovician
Silurian

See also

Related Research Articles

<span class="mw-page-title-main">Trilobite</span> Class of extinct, Paleozoic arthropods

Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.

<span class="mw-page-title-main">Acritarch</span> Microfossils

Acritarchs are organic microfossils, known from approximately 1800 million years ago to the present. The classification is a catch all term used to refer to any organic microfossils that cannot be assigned to other groups. Their diversity reflects major ecological events such as the appearance of predation and the Cambrian explosion.

<span class="mw-page-title-main">Nektaspida</span> Extinct order of arthropods

Nektaspida is an extinct order of non-mineralised artiopodan arthropods. They are known from the mid-Cambrian to the upper Silurian. Originally classified as trilobites, which they superficially resemble, they are now placed as close relatives as members of the Trilobitomorpha within Artiopoda. The order is divided into three major families; Emucarididae, Liwiidae, and Naraoiidae.

<span class="mw-page-title-main">Eurypterid</span> Order of arthropods (fossil)

Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event 251.9 million years ago.

<span class="mw-page-title-main">Cambrian–Ordovician extinction event</span> Mass extinction event about 488 million years ago

The Cambrian–Ordovician extinction event occurred approximately 488 million years ago (mya). This early Phanerozoic Eon extinction event eliminated many brachiopods and conodonts, and severely reduced the number of trilobite species. The Period in the Cambrian extinction in which most of the extinction occurred was the Caerfai Period.

<i>Flexicalymene</i> Genus of trilobites (fossil)

Flexicalymene Shirley, 1936. is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and Family Calymenidae. Flexicalymene specimens can be mistaken for Calymene, Gravicalymene, Diacalymene and a few other Calymenina genera. They are used as an index fossil in the Ordovician. Ohio and North America are particularly known for being rich with Flexicalymene fossils.

<i>Diacalymene</i>

Diacalymene is a genus of trilobite from the order Phacopida, suborder Calymenina. It includes the species D. ouzregui, D. clavicula, D. diademata and D. gabrielsi. It lived in the Ordovician and Silurian periods.

<i>Choia</i> Genus of sponges

Choia is a genus of extinct demosponge ranging from the Cambrian until the Lower Ordovician periods. Fossils of Choia have been found in the Burgess Shale in British Columbia; the Maotianshan shales of China; the Wheeler Shale in Utah; and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott.

<i>Pytine</i> Extinct genus of trilobites

Pytine is an extinct genus of asaphid trilobites. Species lived during the later part of the Arenig stage of the Ordovician Period, approximately 478 to 471 million years ago. Various species are found in the Svalbard, Valhallfonna Formation, Olenidsletta, Member, of Spitzbergen, Norway, the Megistaspis (Paramegistaspis) planilimbata Zone of the 'Shumardia Shale' of Sweden, Jujuy Province, Argentina, early Arenig-aged strata of Jiangxi province, China, and Darriwilian-aged strata in Western Hunan province, China. The type species, P. graia, has seven thorax segments, and lacks the rapier-like glabellar spine, that occurs in many other raphiophorids. The Chinese species, by contrast, have only six thoracic segments. So far, only the type species, and one of the Chinese species, P. laevigata, are known from complete specimens.

<i>Cyphaspis</i> Extinct genus of trilobites

Cyphaspis is a genus of small trilobite that lived from the Late Ordovician to the Late Devonian. Fossils have been found in marine strata in what is now Europe, Africa and North America. Various species had a compact body, and a large, bulbous glabellum. Many species had long spines arranged similarly to closely related genera, such as Otarian, Otarionella, Chamaeleoaspis, and Namuropyge.

<i>Homalonotus</i>

Homalonotus is an extinct genus of trilobites in the order Phacopida. It contains several species, including H. armatus and H. roemeri. It is closely related to other trilobites such as Arduennella and Dipleura..

<i>Lonchodomas</i> Extinct genus of trilobites

Lonchodomas is a genus of trilobites, that lived during the Ordovician. It was eyeless, like all raphiophorids, and had a long straight sword-like frontal spine, that gradually transforms into the relatively long glabella. Both the glabellar spine and the backward directed genal spines are subquadrate in section. Lonchodomas has five thorax segments and the pleural area of the pygidium has two narrow furrows. Lonchodomas occurred in what are today Argentina, Canada (Newfoundland), Estonia, Latvia, Norway, Sweden, the Russian Federation and the United States.

<i>Bumastus</i> Extinct genus of trilobites

Bumastus is an extinct genus of corynexochid trilobites which existed from the Early Ordovician period to the Late Silurian period. They were relatively large trilobites, reaching a length of 6 in (15 cm). They were distinctive for their highly globular, smooth-surfaced exoskeleton. They possessed well-developed, large compound eyes and were believed to have dwelled in shallow-water sediments in life.

<span class="mw-page-title-main">Paleontology in Illinois</span>

Paleontology in Illinois refers to paleontological research occurring within or conducted by people from the U.S. state of Illinois. Scientists have found that Illinois was covered by a sea during the Paleozoic Era. Over time this sea was inhabited by animals including brachiopods, clams, corals, crinoids, sea snails, sponges, and trilobites.

<span class="mw-page-title-main">Paleontology in Wisconsin</span>

Paleontology in Wisconsin refers to paleontological research occurring within or conducted by people from the U.S. state of Wisconsin. The state has fossils from the Precambrian, much of the Paleozoic, and the later part of the Cenozoic. Most of the Paleozoic rocks are marine in origin. Because of the thick blanket of Pleistocene glacial sediment that covers the rock strata in most of the state, Wisconsin’s fossil record is relatively sparse. In spite of this, certain Wisconsin paleontological occurrences provide exceptional insights concerning the history and diversity of life on Earth.

<span class="mw-page-title-main">Raphiophoridae</span> Extinct family of trilobites

Raphiophoridae is a family of small to average-sized trilobites that first occurred at the start of the Ordovician and became extinct at the end of the Middle Silurian.

<i>Furca</i> (genus) Extinct genus of arthropods

Furca is an extinct genus of marrellomorph arthropod known from the Sandbian stage of the Czech Republic, with a single currently described species, Furca bohemica. A tentative additional species, "Furca mauretanica": was proposed for specimens discovered in Morocco, but this species remains a nomen nudum until formally published, and probably belongs in a new separate genus.

<i>Placoparia</i> Genus of trilobites

Placoparia is a genus of trilobites of average size that lived during the late Lower to the early Upper Ordovician on the paleocontinents Gondwana, Avalonia and Laurentia, now the Czech Republic, France, Germany, Morocco, Portugal, Spain and Wales. Its headshield is semi-circular to rectangular with rounded frontal corners. It lacks eyes, but eye ridges are present. The fact that the facial sutures are opisthoparian is an exception in the otherwise proparian Cheirurina. The thorax has 11 or 12 segments, with the axis slightly wider than the ribs to its sides. The tips of the pleurae are free, which resembles an old-fashion central heating radiator. The axis in the small tailshield consists of four rings and a minute endpiece. The four pleurae end in spatulate spines that fit to corresponding indentations in the cephalon.

<i>Bulbaspis</i> Extinct genus of trilobites

Bulbaspis is a late Ordovician genus of asaphid trilobites of the family Raphiophoridae found primarily in upper Ordovician-aged deepwater marine strata of Kazakhstan, China, and possibly Tasmania. Species of Bulbaspis are similar to other raphiophorids such as Ampyx and Raphiophorus, save that the long spine that emanates from the glabella of the latter two genera has been modified into a knob-like or bulb-like structure in Bulbaspis that developed incrementally in the animal's growth. The function of the bulb is currently unknown: one hypothesis suggests sexual selection may have had a role in its evolution in the genus.

<i>Odontopleura</i> Extinct genus of trilobites

Odontopleura is a genus of spinose odontopleurid trilobite in the family Odontopleuridae, and is the type genus of that family and of Odontopleurida. The various species are found in Upper Ordovician to Middle Devonian marine strata throughout the world. The best studied fossils are of the type species, O. ovata, from the Wenlock-aged Liteň Formation in Loděnice, in Bohemia, Czech Republic, and, southeastern Gotland, of Sweden.

References

  1. H H Swinnerton - Outlines of Palaeontology, Edward Arnold (Publishers) Ltd, 1972. ISBN   0-7131-2357-5.
  2. British Palaeozoic Fossils - Natural History Museum 4th edition 1975. Plate 8. ISBN   0-565-05624-7.
  3. Fossil 'conga lines' reveal origins of animal swarms National Geographic, 17 October 2019.
  4. Vannier J, Vidal M, Marchant R, El Hariri K, Kouraiss K, Pittet B, El Albani A, Mazurier A and Martin E (2019) "Collective behaviour in 480-million-year-old trilobite arthropods from Morocco", Nature: Scientific Reports, 9: 14941. doi : 10.1038/s41598-019-51012-3
  5. Ampyx at Fossilworks.org