Asaphida

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Asaphida
Temporal range: Middle Cambrian–Silurian
Asaphus.jpg
Asaphus lepidurus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Subclass: Librostoma
Order: Asaphida
Fortey & Chatterton, 1988
Superfamilies

Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies (Anomocaroidea, Asaphoidea, Cyclopygoidea, Dikelocephaloidea, Remopleuridoidea and Trinucleioidea), but no suborders. Asaphids comprise some 20% of described fossil trilobites.

Contents

In 2020, the superfamily Trinucleioidea was proposed to be raised to an order (Trinucleida) and removed from Asaphida. [1]

Morphology

Asaphellus sp Asaphellus fossil from Mila formation, Ordovician period, Damghan, Iran 01.jpg
Asaphellus sp

The Asaphids generally have cephalon (head) and pygidium (tail) parts similar in size, and most species have a prominent median ventral suture. Heads are often flat, and carapace furrows in the head area are often faint or not visible. Thoracic segments typically number 5 - 12, though some species have as few as two and some as many as 30. They also generally have a wide doublure, or rim, that surrounds the cephalon. This causes some specimens to be described as having a characteristic "snowplow" shaped cephalon. When present, eyes are typically large.

Asaphus kowalewski Trilobite2.jpg
Asaphus kowalewski

One Asaphida line, the Superfamily Asaphoidea, shows a continuous evolution of eyestalks, from individuals with stubby eyes to Asaphus kowalewskii , a trilobite with long eyestalks. This line is found in the Middle Ordovician Asery Level deposits of the Volkhov River region near Saint Petersburg, Russia. During the Ordovician, the region that is now Eastern Europe was a shallow inland sea. This eyestalk development is believed to be an adaptation to changes in turbidity during this time, with eye-stalked trilobites like A. kowalewski presumably arising in a time of increased turbidity. One thought is that this trilobite may have lain in wait for prey buried in the bottom sediment with only its periscope eyestalks protruding.

The major extinction event marking the end of the Ordovician Period reduced the diversity of all trilobite orders with most asaphid families disappearing. The only surviving asaphids were members of superfamily Trinucleioidea, and they too disappeared before the end of the Silurian Period.

Damghanampyx ghobadipour Damghan Ampyx ghobadipour fossil from Lashkark Formation, Ordovician, Damghan, Iran.jpg
Damghanampyx ghobadipour

Taxonomy

The following superfamilies and families are included: [2]

Related Research Articles

<span class="mw-page-title-main">Naraoiidae</span> Family of arthropods (fossil)

Naraoiidae is a family, of extinct, soft-shelled trilobite-like arthropods, that belongs to the order Nektaspida. Species included in the Naraoiidae are known from the second half of the Lower Cambrian to the end of the Upper Silurian. The total number of collection sites is limited and distributed over a vast period of time: Maotianshan Shale and Balang Formation (China), Burgess Shale and Bertie Formation (Canada), the Šárka Formation, Emu Bay Shale (Australia), Idaho and Utah (USA). This is probably due to the rare occurrence of the right circumstances for soft tissue preservation, needed for these non-calcified exoskeletons.

<span class="mw-page-title-main">Ordovician</span> Second period of the Paleozoic Era 485–444 million years ago

The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Ma) to the start of the Silurian Period 443.8 Mya.

<span class="mw-page-title-main">Trilobite</span> Class of extinct, Paleozoic arthropods

Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.

<i>Asaphus kowalewskii</i> Species of trilobite

Asaphus kowalewskii is one of the 35 species of trilobites of the genus Asaphus. Fossils of this species are popular among collectors because of their prominent stalked eyes, many an inch or more in length.

<span class="mw-page-title-main">Ptychopariida</span> Extinct order of trilobites

Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half of the Lower Cambrian, and the last species did not survive the Ordovician–Silurian extinction event.

<span class="mw-page-title-main">Lichida</span> Extinct order of trilobites

Lichida is an order of typically spiny trilobite that lived from the Furongian to the Devonian period. These trilobites usually have 8–13 thoracic segments. Their exoskeletons often have a grainy texture or have wart or spine-like tubercles. Some species are extraordinarily spiny, having spiny thoracic segments that are as long or longer than the entire body, from cephalon (head) to pygidium (tail). The sections of the pygidia are leaf-like in shape and also typically end in spines.

<i>Dalmanites</i> Extinct genus of trilobites

Dalmanites is a genus of trilobite in the order Phacopida. They lived from the Late Ordovician to Middle Devonian.

<span class="mw-page-title-main">Spiriferida</span> Order of brachiopods

Spiriferida is an order of extinct articulate brachiopod fossils which are known for their long hinge-line, which is often the widest part of the shell. In some genera it is greatly elongated, giving them a wing-like appearance. They often have a deep fold down the center of the shell. The feature that gives the spiriferids their name ("spiral-bearers") is the internal support for the lophophore; this brachidium, which is often preserved in fossils, is a thin ribbon of calcite that is typically coiled tightly within the shell.

<i>Isotelus</i> Extinct genus of trilobites

Isotelus is a genus of asaphid trilobites from the middle and upper Ordovician period, fairly common in the Northeastern United States, northwest Manitoba, southwestern Quebec and southeastern Ontario. One species, Isotelus rex, is currently the world's largest trilobite ever found as a complete fossil, and was probably exceeded in size only by Hungioides bohemicus, in which the specimens probably exceeding Isotelus rex in size are only known from partial remains.

<i>Deiphon</i> Genus of trilobites

Deiphon is a distinctive genus of Silurian phacopid trilobites of the family Cheiruridae found in Western and Central Europe, and in Central and Eastern United States. The type species, D. forbesi, from England, Bohemia, and Sweden, was discovered and described by the French paleontologist, Joachim Barrande in 1850.

In the geological timescale, the Llandovery Epoch occurred at the beginning of the Silurian Period. The Llandoverian Epoch follows the massive Ordovician-Silurian extinction events, which led to a large decrease in biodiversity and an opening up of ecosystems.

<i>Aulacopleura</i> Extinct genus of trilobites

Aulacopleura is a genus of proetid trilobite that lived from the Middle Ordovician to the Middle Devonian. Some authors may classify this group as subgenus Otarion (Aulacopleura). The cephalon is semicircular or semielliptical, with border and preglabellar field. The glabella is short, with or without defined eye ridges connecting it with eyes of variable size. Spines at the rear outer corners of the cephalon are present, typically reaching back to the 2nd to 4th thorax segment. The 'palate' is not connected to the dorsal shield of the cephalon. The cephalon is pitted, or has small tubercles. The thorax has up to 22 segments. The pleural ends are usually rounded. The pygidium is small (micropygous), with an even margin. A. koninckii had a modern type of compound eye.

<span class="mw-page-title-main">Asaphidae</span> Extinct family of trilobites


Asaphidae is a family of asaphid trilobites. Although the first genera originate in Upper Cambrian marine strata, the family becomes the most widely distributed and most species-rich trilobite family during the Ordovician. 754 species assigned to 146 genera are included in Asaphidae.

<i>Bumastus</i> Extinct genus of trilobites

Bumastus is an extinct genus of corynexochid trilobites which existed from the Early Ordovician period to the Late Silurian period. They were relatively large trilobites, reaching a length of 6 in (15 cm). They were distinctive for their highly globular, smooth-surfaced exoskeleton. They possessed well-developed, large compound eyes and were believed to have dwelled in shallow-water sediments in life.

Fossils of many types of water-dwelling animals from the Devonian period are found in deposits in the U.S. state of Michigan. Among the more commonly occurring specimens are bryozoans, corals, crinoids, and brachiopods. Also found, but not so commonly, are armored fish called placoderms, snails, sharks, stromatolites, trilobites and blastoids.

<span class="mw-page-title-main">Eodiscina</span>

Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.

<i>Ogygiocarella</i> Extinct genus of trilobites

Ogygiocarella Brongniart, 1822, is a genus of asaphid trilobites. It occurred during the Middle Ordovician.

<span class="mw-page-title-main">Cyclopygidae</span> Extinct family of trilobites

Cyclopygidae is a family of asaphid trilobites from the Ordovician. Cyclopygids had an extratropical distribution, and there is evidence that they lived in darker parts of the water column. Cyclopygids are characterized by enlarged eyes, with a wide angle of view, both horizontal and vertical, reminiscent of the eyes of dragonflies. These typically touch the glabella directly on the side. Cyclopygids all lack genal spines, but Symphysops carries a forward directed frontal spine on the glabella. It is presumed that at least the members of the genus Pricyclopyge swam upside down and had bioluminescent organs on the third thorax segment. Cyclopygids had between 7 and 5 thorax segments, a wide and stout axis, and short side lobes.

<span class="mw-page-title-main">Raphiophoridae</span> Extinct family of trilobites

Raphiophoridae is a family of small to average-sized trilobites that first occurred at the start of the Ordovician and became extinct at the end of the Middle Silurian.

<span class="mw-page-title-main">Librostoma</span> Extinct subclass of trilobites

Librostoma is a subclass of trilobites defined by having a natant hypostome, which is a hypostome that is free from the anterior doublure and aligned with the anterior of the glabella, this is unlike a conterminant hypostome, which is attached to the exoskeleton.

References

  1. Bignon, Arnaud; Waisfeld, Beatriz G.; Vaccari, N. Emilio; Chatterton, Brian D. E. (2020-07-02). "Reassessment of the Order Trinucleida (Trilobita)". Journal of Systematic Palaeontology. 18 (13): 1061–1077. Bibcode:2020JSPal..18.1061B. doi:10.1080/14772019.2020.1720324. ISSN   1477-2019. S2CID   212995185.
  2. S. M. Gon III. "Order Asaphida" . Retrieved 9 July 2018.