| Entomaspididae | |
|---|---|
 | |
| Entomaspis radiata | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | † Trilobita |
| Order: | † Harpetida |
| Family: | † Entomaspididae Ulrich, 1931 |
| Genera | |
Entomaspididae is a family of harpetid trilobites that ranges from the Upper Cambrian to Lower Ordovician of marine strata in China and the United States. [1]
Entomaspidids, as in other harpetid trilobites, have proportionally large, vaulted cephalons, shaped like croissants or bonnets, with the cheeks freed to be modified into long and curved spines in Entomaspis and Hypothetica . Although the eyes of Entomaspididae are small compared to many other trilobites, the eyes are large when compared to other harpetids bearing eyes. Entomaspidids can be further distinguished from other harpetids by having a continuous border along the rim of the pygidium composed of small, stub-like spines. [2] In overall appearance, the typical Entomaspidid is a near-circular, medallion-like trilobite.
Entomaspididae is thought to represent the transition between the ancestral Ptychopariida, and the harpetids. Many researchers question the validity of Entompaspididae as a distinct family, and merge its genera into Harpetidae. [3]
Ulrich erected "Entomaspididae" to contain species of Entomaspis and Hypothetica , from the Latest Cambrian or Earliest Ordovician of Utah. [1]
The first entomaspidids appear in the fossil record near the close of the Cambrian period in marine strata of Utah and China. They persist into the Early Ordovician before disappearing from the fossil record.
The three genera traditionally placed in Entomaspididae include the type genus, Entomaspis , whose species are found in Utah and Missouri, and Hypothetica , which lived sympatrically with Utah species of Entomaspis, and Baikadamaspis , of Late Cambrian marine strata in China. Many authorities merge Hypothetica into Entomaspis .
Trilobites are a group of extinct marine artiopodan arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period, and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 300 million years.
Harpetida is one of the eleven orders of the extinct arthropod class Trilobita. The first harpetid trilobites appear in the Upper Cambrian, and the last species die out at the end of the Devonian period.
Phacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian, with a broader time range described from the Late Ordovician. It was a rounded animal, with a globose head and large eyes, and probably fed on detritus. Phacops is often found rolled up ("volvation"), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus.
Olenoides was a trilobite from the Cambrian period. Its fossils are found well-preserved in the Burgess Shale in Canada. It grew up to 10 cm long.
Naraoia is a genus of small to average size marine arthropods within the family Naraoiidae, that lived from the early Cambrian to the late Silurian period. The species are characterized by a large alimentary system and sideways oriented antennas.
Asaphus is a genus of trilobites that is known from the Lower and Middle Ordovician of northwestern Europe.
Pytine is an extinct genus of asaphid trilobites. Species lived during the later part of the Arenig stage of the Ordovician Period, approximately 478 to 471 million years ago. Various species are found in the Svalbard, Valhallfonna Formation, Olenidsletta, Member, of Spitzbergen, Norway, the Megistaspis (Paramegistaspis) planilimbata Zone of the 'Shumardia Shale' of Sweden, Jujuy Province, Argentina, early Arenig-aged strata of Jiangxi province, China, and Darriwilian-aged strata in Western Hunan province, China. The type species, P. graia, has seven thorax segments, and lacks the rapier-like glabellar spine, that occurs in many other raphiophorids. The Chinese species, by contrast, have only six thoracic segments. So far, only the type species, and one of the Chinese species, P. laevigata, are known from complete specimens.
Yunnanocephalus is a genus of ptychopariid trilobite. It lived during the late Atdabanian and Botomian stages, in what are currently Antarctica, Australia and China. It was a "moderately common" member of the Chengjiang Fauna. Yunnanocephalus is the only genus currently assigned to the Yunnanocephalidae family.
Crepicephalus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period.
Meteoraspis is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae. The various species lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Meteoraspis are characteristic of Late Cambrian strata in North America, though they are found in Late Cambrian strata elsewhere in the world, such as M. nevensis from Victoria Land, Antarctica.
Asaphiscus is a genus of trilobite that lived in the Cambrian. Its remains have been found in Australia and North America, especially in Utah.
Ptychoparia is a genus of ptychopariid trilobites, and is the type genus of the family Ptychopariidae, and the order Ptychopariida.
Pricyclopyge is a genus of trilobites assigned to the family Cyclopygidae that occurs throughout the Ordovician. Pricyclopyge had an extratropical distribution, and there is evidence that it lived in darker parts of the water column. Pricyclopyge has huge eyes, an inverted pear-shaped glabella, six thorax segments, with on the 3rd two small discs. Pricyclopyge is known from what are today China, the Czech Republic, France, and the United Kingdom.
Carolinites is a genus of trilobite, assigned to the Telephinidae family, that occurs during the Lower and Middle Ordovician. Carolinites had a pantropical distribution, and there is evidence that it lived in upper parts of the water column. The free cheeks of Carolinites are largely covered by its huge eyes, except for the attachment of large genal spines that extend downward, backward and lateral and gradually curving further backward. The glabella is slightly bulbous, the occipital ring is well defined, but further transglabellar furrows are lacking. The thorax has 10 segments. The axis of the pygidium is highly vaulted, with a curved spine emerging almost perpendicular to the midline and ending parallel to it and a node on each of the other three segments. Carolinites is known from what are today Australia (Tasmania), Canada (Alberta), China, France, Spitsbergen, and the United States (Utah).
Angelina Salter, 1859, is a genus of ptychopariid trilobite belonging to the Family Olenidae, Suborder olenina. It lived during the Tremadocian Stage, lowermost of the two standard worldwide divisions forming the Lower Ordovician Series and lowest of the seven stages within the Ordovician System. It encompasses all rocks formed during Tremadocian times, which spanned the interval between 485.4 million and 477.7 million years ago. Fossilized remains of Angelina are known from Wales, Central and South America. It differs from most other Triarthrinae in being larger, with a relatively narrow glabella, the occipital ring poorly defined, and lateral glabellar furrows relatively obscure. Eyes are placed midlength that of the cephalon and the facial sutures converge on the front border at the midline. Species also have long genal spines.
Orygmaspis is a genus of asaphid trilobite with an inverted egg-shaped outline, a wide headshield, small eyes, long genal spines, 12 spined thorax segments and a small, short tailshield, with four pairs of spines. It lived during the Upper Cambrian in what are today Canada and the United States.
Genevievella is a genus of trilobites with a short inverted egg-shaped outline, a wide headshield, small eyes, and long genal spines. The backrim of the headshield is inflated and overhangs the first of the 9 thorax segments. The 8th thorax segment from the front bears a backward directed spine that reaches beyond the back end of the exoskeleton. It has an almost oval tailshield with 5 pairs of pleural furrows. It lived during the Upper Cambrian in what are today Canada and the United States.
Placoparia is a genus of trilobites of average size that lived during the late Lower to the early Upper Ordovician on the paleocontinents Gondwana, Avalonia and Laurentia, now the Czech Republic, France, Germany, Morocco, Portugal, Spain and Wales. Its headshield is semi-circular to rectangular with rounded frontal corners. It lacks eyes, but eye ridges are present. The fact that the facial sutures are opisthoparian is an exception in the otherwise proparian Cheirurina. The thorax has 11 or 12 segments, with the axis slightly wider than the ribs to its sides. The tips of the pleurae are free, which resembles an old-fashion central heating radiator. The axis in the small tailshield consists of four rings and a minute endpiece. The four pleurae end in spatulate spines that fit to corresponding indentations in the cephalon.
Tricrepicephalus is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae with species of average size. Its species lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Tricrepicephalus are widespread in Late Cambrian deposits in North America, but is also known from one location in South-America. Tricrepicephalus has an inverted egg-shaped exoskeleton, with three characteristic pits in the fold that parallels the margin of the headshield just in front of the central raised area. The articulating middle part of the body has 12 segments and the tailshield carries two long, tubular, curved pygidial spines that are reminiscent of earwig's pincers that rise backwards from the plain of the body at approximately 30°.
Entomaspis is an extinct genus of harpetid trilobite from Upper Cambrian to Early Ordovician marine strata of the United States. Species are typified by their proportionally large, vaulted, croissant-shaped or bonnet-shaped cephalons that have the cheeks freed to become elongated, curved librigenial spines, and by their comparatively large, crescent-shaped eyes.