Artiodactyls are placental mammals belonging to the order Artiodactyla. Typically, they are ungulates which bear weight equally on two of their five toes: the third and fourth, often in the form of a hoof. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, most perissodactyls bear weight on an odd number of the five toes. Another difference between the two is that many artiodactyls digest plant cellulose in one or more stomach chambers rather than in their intestine as perissodactyls do. The advent of molecular biology, along with new fossil discoveries, found that cetaceans fall within this taxonomic branch, being most closely related to hippopotamuses. Some modern taxonomists thus apply the name Cetartiodactyla to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use "even-toed ungulates" to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature.
Hippopotamidae is a family of stout, naked-skinned, and semiaquatic artiodactyl mammals, possessing three-chambered stomachs and walking on four toes on each foot. While they resemble pigs physiologically, their closest living relatives are the cetaceans. They are sometimes referred to as hippopotamids.
Anthracotherium is an extinct genus of artiodactyls characterized by having 44 teeth, with five semi-crescentic cusps on the crowns of the upper molars. The genus ranged from the middle Eocene period until the early Miocene, having a distribution throughout Eurasia. Material subjectively assigned to Anthracotherium from Pakistan suggests the last species died out soon after the start of the Miocene.
Megistotherium is an extinct genus of hyaenodont belonging to the family Hyainailouridae that lived in Africa.
Pan Am Flight 103 (PA103/PAA103) was a regularly scheduled Pan Am transatlantic flight from Frankfurt to Detroit via a stopover in London and another in New York City. The transatlantic leg of the route was operated by Clipper Maid of the Seas, a Boeing 747 registered N739PA. Shortly after 19:00 on 21 December 1988, while the aircraft was in flight over the Scottish town of Lockerbie, it was destroyed by a bomb, killing all 243 passengers and 16 crew in what became known as the Lockerbie bombing. Large sections of the aircraft crashed in a residential street in Lockerbie, killing 11 residents. With a total of 270 fatalities, it is the deadliest terrorist attack in the history of the United Kingdom.
Anthracotheriidae is a paraphyletic family of extinct, hippopotamus-like artiodactyl ungulates related to hippopotamuses and whales. The oldest genus, Elomeryx, first appeared during the middle Eocene in Asia. They thrived in Africa and Eurasia, with a few species ultimately entering North America during the Oligocene. They died out in Europe and Africa during the Miocene, possibly due to a combination of climatic changes and competition with other artiodactyls, including pigs and true hippopotamuses. The youngest genus, Merycopotamus, died out in Asia during the late Pliocene, possibly for the same reasons. The family is named after the first genus discovered, Anthracotherium, which means "coal beast", as the first fossils of it were found in Paleogene-aged coal beds in France. Fossil remains of the anthracothere genus were discovered by the Harvard University and Geological Survey of Pakistan joint research project (Y-GSP) in the well-dated middle and late Miocene deposits of the Pothohar Plateau in northern Pakistan.
Merycopotamus is an extinct genus of Asian anthracothere that appeared during the Middle Miocene, and died out in the Late Pliocene. At the height of the genus' influence, species ranged throughout southern Asia. With the extinction of the last species, M. dissimilis, the lineage of anthracotheres came to an end. Merycopotamus was closely related to the anthracothere genus Libycosaurus, which, unlike the former, never left Africa. In fact, some African fossils originally placed in Merycopotamus, but are now referred to Libycosaurus.
Anthracothema was a genus of extinct artiodactyl ungulate mammals that lived in Myanmar during the late Eocene.
Paraentelodon is an extinct entelodont from the Late Oligocene and Oligocene-Miocene boundary of Asia. The fossils of the type species P. intermedium were found in Georgia, Kazakhstan and China. An indeterminate species represents in Bugti Hills which is the late Oligocene of Pakistan.
Euthecodon is an extinct genus of long-snouted crocodile. It was common throughout much of Africa during the Neogene, with fossils being especially common in Kenya, Ethiopia, and Libya. Although superficially resembling that of gharials, the long snout was a trait developed independently from that of other crocodilians and suggests a diet of primarily fish. Euthecodon coexisted with a wide range of other crocodiles in the areas it inhabited before eventually going extinct during the Pleistocene.
Allaeochelys is an extinct genus of Carettochelyid turtle, known from the Eocene to Miocene of Europe, Asia, North America and Africa.
Rimasuchus is an extinct genus of crocodile from the Miocene of Egypt and possibly Libya. Only one species - Rimasuchus lloydi - is currently known. It was previously thought to be a species of Crocodylus, but is now thought to be more closely related to the modern African dwarf crocodiles (Osteolaemus).
Jaggermeryx is an extinct genus of semiaquatic anthracothere, ungulates related to hippopotamuses, from the Early Miocene Moghara Formation in Egypt. The genus was named after Mick Jagger.
Bugtitherium is an extinct genus of anthracothere found in late Oligocene (Chattian) deposits in the Bugti Hills of Baluchistan, Pakistan.
The geology of Libya formed on top of deep and poorly understood Precambrian igneous and metamorphic crystalline basement rock. Most of the country is intra-craton basins, filled with thick layers of sediment. The region experienced long-running subsidence and terrestrial sedimentation during the Paleozoic, followed by phases of volcanism and intense folding in some areas, and widespread flooding in the Mesozoic and Cenozoic due to a long marine transgression. Libya has the largest hydrocarbon reserves in Africa, as well as deposits of evaporites.
Microbunodon was a genus of extinct artiodactyl mammals in the family Anthracotheriidae. It lived between the upper Eocene and the lower Pliocene. Its fossil remains have been found in Europe and Asia.
Brachyodus was a genus of anthracothere that lived in Europe during the Early Miocene.
Afrocygnus is an extinct genus of swan, which lived during the Late Miocene, and perhaps up to the Late Pliocene, in what is today North Africa. The only genus of swan known in Africa, aside from fragmentary Pleistocene remains found in East Africa and from occasional observations of vagrant European swans along the Mediterranean coast, it lived in what was during the Miocene a damp wetland spanning from Libya to Chad, alongside the Antracothere Libycosaurus and the early Homininae Sahelanthropus. The genus is considered as the sister taxon of the extant genus Cygnus. Fossils of the genus have been uncovered in the Sahabi Formation of Cyrenaica in Libya, and in the Toros-Menalla locality in the Djurab Desert of Northern Chad.
The microbunodontines were an extinct subfamily of anthracotheres that were predominately a Paleogene group of Eurasian artiodactyls. The group died out at the end of the Late Miocene. It comprised the genera Anthracokeryx, Geniokeryx, Microbunodon, and possibly Etruscotherium. They are different from the other anthracothere lineages by their smaller size, slenderer limbs and male specimens having laterally compressed, longer canines. They were originally classified as members of the other subfamily of anthracotheres, Anthracotheriinae but recent phylogenetic studies have found them to be their own clade sister to Bothriodontinae.
Ancodonta is an infraorder of semiaquatic artiodactyl ungulates including modern hippopotamus and all mammals closer to hippos than to cetaceans (whales). Ancodonts first appeared in the Middle Eocene, with some of the earliest representatives found in fossil deposits in Southeast Asia. Throughout their evolutionary history they have occupied different browsing and grazing niches in North America, Eurasia and Africa. The last continent is notable as they were among the first laurasiatherian mammals to have migrated to Africa from Europe, where they competed with the native afrothere herbivores for the same niches. Of the nearly 50 genera that have existed, only two of them are extant – Choeropsis and Hippopotamus. The interrelationships within the ancodonts has been contended. The traditional notion is that there at minimum two families Anthracotheriidae and Hippopotamidae and were merely sister taxa. However many detailed research of the dentition among ancodonts, as well as how some anthracotheres were similar to hippos in appearance, lead the current consensus where Anthracotheriidae is paraphyletic to Hippopotamidae. Among the anthracotheres members of Bothriodontinae are among the closest to the ancestry of hippos, with the Oligocene aged Epirigenys from Lokon, Turkana, Kenya being the sister taxon to hippos. In response of this many similar clade names have been used for this clade.
