The Pennsylvanian is, on the ICS geologic timescale, the younger of two subperiods of the Carboniferous Period. It lasted from roughly 323.2 million years ago to 298.9 million years ago. As with most other geochronologic units, the rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal beds of this age are widespread.
Odonata is an order of predatory flying insects that includes the dragonflies and damselflies. The two groups are distinguished with dragonflies usually being bulkier with large compound eyes together and wings spread up or out at rest, while damselflies are usually more slender with eyes placed apart and wings folded together along body at rest. Adult odonates can land and perch, but rarely walk.
Meganeura is a genus of extinct insects from the Late Carboniferous. They resembled and are related to the present-day dragonflies and damselflies, and were predatory, with their diet mainly consisting of other insects. The genus belongs to the Meganeuridae, a family including other similarly giant dragonfly-like insects ranging from the Late Carboniferous to Middle Permian. With a wingspan about 65–75 cm (2.13–2.46 ft), M. monyi is one of the largest-known flying insect species.
Ensifera is a suborder of insects that includes the various types of crickets and their allies including: true crickets, camel crickets, bush crickets or katydids, grigs, weta and Cooloola monsters. This and the suborder Caelifera make up the order Orthoptera. Ensifera is believed to be a more ancient group than Caelifera, with its origins in the Carboniferous period, the split having occurred at the end of the Permian period. Unlike the Caelifera, the Ensifera contain numerous members that are partially carnivorous, feeding on other insects, as well as plants.
Meganisoptera is an extinct order of large dragonfly-like insects, informally known as griffenflies or (incorrectly) as giant dragonflies. The order was formerly named Protodonata, the "proto-Odonata", for their similar appearance and supposed relation to modern Odonata. They range in Palaeozoic times. Though most were only slightly larger than modern dragonflies, the order includes the largest known insect species, such as the late Carboniferous Meganeura monyi and the even larger early Permian Meganeuropsis permiana, with wingspans of up to 71 centimetres (28 in).
The most recent understanding of the evolution of insects is based on studies of the following branches of science: molecular biology, insect morphology, paleontology, insect taxonomy, evolution, embryology, bioinformatics and scientific computing. It is estimated that the class of insects originated on Earth about 480 million years ago, in the Ordovician, at about the same time terrestrial plants appeared. Insects are thought to have evolved from a group of crustaceans. The first insects were landbound, but about 400 million years ago in the Devonian period one lineage of insects evolved flight, the first animals to do so. The oldest insect fossil has been proposed to be Rhyniognatha hirsti, estimated to be 400 million years old, but the insect identity of the fossil has been contested. Global climate conditions changed several times during the history of Earth, and along with it the diversity of insects. The Pterygotes underwent a major radiation in the Carboniferous while the Endopterygota underwent another major radiation in the Permian.
Haptodus is an extinct genus of basal sphenacodont, member of the clade that includes therapsids and hence, mammals. It was at least 1.5 metres (5 ft) in length. It lived in present-day France during the Early Permian. It was a medium-sized predator, feeding on insects and small vertebrates.
Namurotypus is an extinct genus of griffinfly with a single described species Namurotypus sippeli. It inhabited the large swamps of the Carboniferous period. Namaurotypus had a 15 cm long forewing and did not have secondary male sex organs as in modern dragonflies.
The Odonatoptera are a superorder of ancient winged insects, placed in the probably paraphyletic group Palaeoptera. The dragonflies and damselflies are the only living members of this group, which was far more diverse in the late Paleozoic and contained gigantic species, including the griffinflies of the order Meganisoptera. This lineage dates back at least to the Bashkirian, not quite 320 million years ago.
Meganeuropsis is an extinct genus of griffinfly, order Meganisoptera, known from the Early Permian Wellington Formation of North America, and represents the largest known insect of all time. Meganeuropsis existed during the Artinskian age of the Permian period, 290.1–283.5 mya. The genus includes two described species by Frank Morton Carpenter, fossil insect curator at the Museum of Comparative Zoology at Harvard University:
The Carboniferous rainforest collapse (CRC) was a minor extinction event that occurred around 305 million years ago in the Carboniferous period. The event occurred at the end of the Moscovian and continued into the early Kasimovian stages of the Pennsylvanian.
Bohemiatupus is an extinct genus of griffinfly in the family Meganeuridae and containing a single species Bohemiatupus elegans. The species is known only from the Late Carboniferous, Bolsovian stage, Kladno Formation near the village of Radnice in the Radnice Basin, Czech Republic.
Bechala is a genus of insects that existed during the late Carboniferous in what is now Germany. It was first described by Jan-Michael Ilger and Carsten Brauckmann in 2012, and the type species is B. sommeri. Its type specimen was a wing discovered at the Küchenberg quarry, in the Ziegelschiefer Formation. The wing measurements are 39×6 millimetres. Bechala was originally assigned to the extinct order Megasecoptera, but a restudy instead assigned it to the superorder Odonatoptera.
The Tarsophlebiidae is an extinct family of medium-sized fossil odonates from the Upper Jurassic and Lower Cretaceous period of Eurasia. They are either the most basal member of the damsel-dragonfly grade ("anisozygopteres") within the stem group of Anisoptera, or the sister group of all Recent odonates. They are characterized by the basally open discoidal cell in both pairs of wings, very long legs, paddle-shaped male cerci, and a hypertrophied ovipositor in females.
Paleontology in Oklahoma refers to paleontological research occurring within or conducted by people from the U.S. state of Oklahoma. Oklahoma has a rich fossil record spanning all three eras of the Phanerozoic Eon. Oklahoma is the best source of Pennsylvanian fossils in the United States due to having an exceptionally complete geologic record of the epoch. From the Cambrian to the Devonian, all of Oklahoma was covered by a sea that would come to be home to creatures like brachiopods, bryozoans, graptolites and trilobites. During the Carboniferous, an expanse of coastal deltaic swamps formed in areas of the state where early tetrapods would leave behind footprints that would later fossilize. The sea withdrew altogether during the Permian period. Oklahoma was home a variety of insects as well as early amphibians and reptiles. Oklahoma stayed dry for most of the Mesozoic. During the Late Triassic, carnivorous dinosaurs left behind footprints that would later fossilize. During the Cretaceous, however, the state was mostly covered by the Western Interior Seaway, which was home to huge ammonites and other marine invertebrates. During the Cenozoic, Oklahoma became home to creatures like bison, camels, creodonts, and horses. During the Ice Age, the state was home to mammoths and mastodons. Local Native Americans are known to have used fossils for medicinal purposes. The Jurassic dinosaur Saurophaganax maximus is the Oklahoma state fossil.
Meganeuridae is an extinct family of griffinfly in the order Meganisoptera. There are more than 20 genera and 50 described species in Meganeuridae. This family of flies contain the genus Meganeuropsis, which has one of the biggest animals in the world, Meganeuropsis permiana. This species of fly had the longest insect wing ever found.
Paralogidae is an extinct family of griffinfly in the order Meganisoptera. There are at least two genera and three described species in Paralogidae.
The Bajo de Véliz Formation is stratigraphic formation of the Paleozoic located in the Paganzo Basin, Argentina. This formation lies in the northwestern sector of San Luis, about 25 km west of Santa Rosa de Conlara. Containing a slice of the Carboniferous-Permian transition, it is one of the key upper Paleozoic successions characterizing that era in westernmost Gondwana, which South America was a part of in the Paleozoic.
Permopsocida is an extinct order of insects known from the Carboniferous (Moscovian) to the Mid-Cretaceous. It is part of Paraneoptera, alongside bark lice, bugs and thrips. Within Paraneoptera it is considered to be closer to the clade containing bugs and thrips rather than bark lice, with an estimated divergence during the Late Carboniferous. The group was first named as a suborder by Robert John Tillyard in 1926, and was raised to a full order by Huang et al. in 2016. It is currently divided up into three families, Psocidiidae which is known from the Permian to Liassic. Permopsocidae which is only known from the Permian, and Archipsyllidae, which is known from the Late Triassic to mid-Cretaceous (Cenomanian). While most members of the group are known from compression fossils, several members of Archipsyllidae are 3 dimensionally preserved in Burmese amber, which has helped clarify the morphology and phylogenetic position of the group. The morphology of the mouthparts suggests that they were capable of suction feeding and chewing, with preserved angiosperm pollen grains in the gut of Psocorrhyncha suggesting that at least some members of the group were pollenivorous.
Bojophlebia is an extinct genus of winged insect from the Pennsylvanian period of the Czech Republic. It includes only a single species, Bojophlebia prokopi, and is the only member of the family Bojophlebiidae. Bojophlebia prokopi was first described in 1985 by Jarmila Kukalová-Peck, who originally described it as a large mayfly-like insect. This original interpretation has since been rejected. Most recently, B. prokopi has been treated as a member of the infraclass Hydropalaeoptera, which also includes the Odonatoptera and Panephemeroptera. Bojophlebia is considered a sister group of all other members of the Hydropalaeoptera. A fossil that was described as a nymph of Bojophlebia is now considered to be a separate taxon, Carbotriplura kukalovae. The original description interpreted structures such as eyes and antennae, however these structures cannot be confirmed after restudy, although this may be an example of over-interpretation by Kukalová-Peck, as has happened with other extinct insects such as Carbotriplura and Gerarus.
