Pelmatozoa was once a clade of Phylum Echinodermata. It included stalked and sedentary echinoderms. The main class of Pelmatozoa were the Crinoidea which includes sea lily and feather star.
Pelmetazoa is no longer a classification of Echinodermata. [1]
Pelmetazoa Temporal range: Middle Cambrian – Recent | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Clade: | Ambulacraria |
Phylum: | Echinodermata |
Groups included | |
Pelmatozoa s.l. ( sensu lato ) or Crinozoa s.l., were in some systems a large taxon, sometimes considered a subphylum of echinoderms.
The majority of Pelmatozoa s.l. consisted of the extinct taxon Blastozoa.
Pelmatozoa s.s. ( sensu stricto ) sometimes only include Crinoidea.
The body of these ancient Echinodermates is anchored to the bottom through a stem analogous to the epistome of Phoronidea. They're sessile, only rarely capable of movement. Fossils date back at least to Cambrium so they've been on Earth for over 500 million years.
In the past, they were a numerous group of marine fauna and many groups belong to lead fossils, however today only one smaller class remains extant.
Pelmatozoa s.l. = Crinozoa s.l.:
Formerly even Carpoidea/Homalozoa belonged to Pelmatozoa.
Sources for this chapter: [5] [6] [7] [8] [9] [10] [11] [12] [13] [14] [15] [16] [17] [18] [19] [20] [21] [22] [23] [24] [25] [26] [27] [28] [29] [30] [31] [32] [33] [34] [35] [36] [37] [38] [39] [40] [41] [42] [43] [44] [45] [46] [47] [48] [49]
Crinoids are marine invertebrates that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their juvenile form are commonly called sea lilies, while the unstalked forms, called feather stars or comatulids, are members of the largest crinoid order, Comatulida. Crinoids are echinoderms in the phylum Echinodermata, which also includes the starfish, brittle stars, sea urchins and sea cucumbers. They live in both shallow water and in depths as great as 9,000 meters (30,000 ft).
Edrioasteroidea is an extinct class of echinoderms. The living animal would have resembled a pentamerously symmetrical disc or cushion. They were obligate encrusters and attached themselves to inorganic or biologic hard substrates. A 507 million years old species, Totiglobus spencensis, is actually the first known echinoderm adapted to live on a hard surface after the soft microbial mats that covered the seafloor were destroyed in the Cambrian substrate revolution.
The stylophorans are an extinct, possibly polyphyletic group allied to the Paleozoic Era echinoderms, comprising the prehistoric cornutes and mitrates. It is synonymous with the subphylum Calcichordata. Their unusual appearances have led to a variety of very different reconstructions of their anatomy, how they lived, and their relationships to other organisms.
Blastozoa is a subphylum of extinct Echinoderms characterized by the presence of specialized respiratory structures and brachiole plates used for feeding. It ranged from the Cambrian to the Permian.
Crinozoa is a subphylum of mostly sessile echinoderms, of which the crinoids, or sea lilies and feather stars, are the only extant members. Crinozoans have an extremely extensive fossil history, which may or may not extend into the Precambrian.
The taxonomy of commonly fossilized invertebrates combines both traditional and modern paleozoological terminology. This article compiles various invertebrate taxa in the fossil record, ranging from protists to arthropods. This includes groups that are significant in paleontological contexts, abundant in the fossil record, or have a high proportion of extinct species. Special notations are explained below:
The Eocrinoidea are an extinct class of echinoderms that lived between the Early Cambrian and Late Silurian periods. They are the earliest known group of stalked, arm-bearing echinoderms, and were the most common echinoderms during the Cambrian.
Deuterostomes are bilaterian animals of the superphylum Deuterostomia, typically characterized by their anus forming before the mouth during embryonic development. Deuterostomia is further divided into 4 phyla: Chordata, Echinodermata, Hemichordata, and the extinct Vetulicolia known from Cambrian fossils. The extinct clade Cambroernida is also thought to be a member of Deuterostomia.
Echmatocrinus is a Cambrian animal which resembles a crinoid or an octocoral. Its exact taxonomy is still a subject of debate. It is known only from the Burgess shale. Around 20 specimens of Echmatocrinus are known; these comprise < 0.02% of the community.
Paracrinoidea is an extinct class of blastozoan echinoderms. They lived in shallow seas during the Early Ordovician through the Early Silurian. While blastozoans are usually characterized by types of respiratory structures present, it is not clear what types of respiratory structures paracrinoids likely had. Despite the taxon's name, the paracrinoids are not closely related to crinoids.
Homalozoa is an obsolete extinct subphylum of Paleozoic era echinoderms, prehistoric marine invertebrates. They are also referred to as carpoids.
The Fezouata Formation or Fezouata Shale is a geological formation in Morocco which dates to the Early Ordovician. It was deposited in a marine environment, and is known for its exceptionally preserved fossils, filling an important preservational window beyond the earlier and more common Cambrian Burgess shale-type deposits. The fauna of this geological unit is often described as the Fezouata biota, and the particular strata within the formation which exhibit exceptional preservation are generally termed the Fezouata Lagerstätte.
Rhenopyrgus is an extinct echinoderm in the class Edrioasteroidea, which existed during the Devonian in what is now France and Germany, the Ordovician in Iowa and Illinois, U.S.A.; and the Silurian of Argentina. It was described by Dehm in 1961, and the type species is R. coronaeformis, which was originally described by J. Rievers as a species in the genus Pyrgocystis, in 1961. A new species, R. piojoensis, was described by Colin D. Sumrall, Susana Heredia, Cecilia M. Rodríguez and Ana I. Mestre in 2012, from 116 specimens collected from the Los Espejos Formation in the Loma de Los Piojos locality near San José de Jáchal, Argentina. The species epithet refers to the locality where the specimens were collected from. In 2019 another species, R. viviani, was described by researchers from the Natural History Museum, London, led by Tim Ewin.
The Verulam Formation is a geologic formation and Lagerstätte in Ontario, Canada. It preserves fossils dating back to the Katian stage of the Ordovician period, or Shermanian to Chatfieldian in the regional stratigraphy.
Diploporita is an extinct class of blastozoan that ranged from the Ordovician to the Devonian. These echinoderms are identified by a specialized respiratory structure, called diplopores. Diplopores are a double pore system that sit within a depression on a single thecal (body) plate; each plate can contain numerous diplopore pairs.
Soluta is an extinct class of echinoderms that lived from the Middle Cambrian to the Early Devonian. The class is also known by its junior synonym Homoiostelea. Soluta is one of the four "carpoid" classes, alongside Ctenocystoidea, Cincta, and Stylophora, which made up the obsolete subphylum Homalozoa. Solutes were asymmetric animals with a stereom skeleton and two appendages, an arm extending anteriorly and a posterior appendage called a homoiostele.
Ctenocystoidea is an extinct clade of echinoderms, which lived during the Cambrian and Ordovician periods. Unlike other echinoderms, ctenocystoids had bilateral symmetry, or were only very slightly asymmetrical. They are believed to be one of the earliest-diverging branches of echinoderms, with their bilateral symmetry a trait shared with other deuterostomes. Ctenocystoids were once classified in the taxon Homalozoa, also known as Carpoidea, alongside cinctans, solutes, and stylophorans. Homalozoa is now recognized as a polyphyletic group of echinoderms without radial symmetry. Ctenocystoids were geographically widespread during the Middle Cambrian, with one species surviving into the Late Ordovician.
Yanjiahella biscarpa is an extinct species of Early Cambrian deuterostome which may represent the earliest stem group echinoderm.
Yorkicystis is a genus of edrioasteroid echinoderm that lived 510 million years ago in the Cambrian aged Kinzers Formation in what is now Pennsylvania. This genus is important as it provides some of the oldest evidence of echinoderms losing their hard mineralized outer skeletons. Yorkicystis also shows that some echinoderms lost their skeletons during the Cambrian, which is a greatly different time as to when most other species lost theirs.
Formerly treated as a formal unit of classification with the rank of subphylum, pelmatozoans are now realized to be a heterogeneous assemblage of forms with similar habits but dissimilar ancestry...