Redoubtia | |
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R. polypodia reconstructed as a holothurian-like animal | |
Scientific classification | |
Genus: | † Redoubtia |
Species: | †R. polypodia |
Binomial name | |
Redoubtia polypodia Walcott 1918 | |
Redoubtia polypodia is a fossil from the Burgess Shale that was originally described (and later refigured) by Charles Walcott as a holothurian echinoderm. [1] [2] Its affinity is unclear, though; it has also been compared to lobopodians. [3]
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At 508 million years old, it is one of the earliest fossil beds containing soft-part imprints.
Echinoderm is the common name given to any member of the phylum Echinodermata of marine animals. The adults are recognizable by their radial symmetry, and include such well-known animals as starfish, sea urchins, sand dollars, and sea cucumbers, as well as the sea lilies or "stone lilies". Echinoderms are found at every ocean depth, from the intertidal zone to the abyssal zone. The phylum contains about 7000 living species, making it the second-largest grouping of deuterostomes, after the chordates. Echinoderms are also the largest phylum that has no freshwater or terrestrial (land-based) representatives.
The lobopodians, members of the informal group Lobopodia are worm-like taxa with stubby legs.
Charles Doolittle Walcott was an American paleontologist, administrator of the Smithsonian Institution from 1907 to 1927, and geologist. He is famous for his discovery in 1909 of well-preserved fossils, including some of the oldest soft-part imprints, in the Burgess Shale of British Columbia, Canada. He was described by Stephen Jay Gould as "the finest symbol that I have ever ncountered for the embodiment of conventional beliefs."
Field is an unincorporated community of approximately 169 people located in the Kicking Horse River valley of southeastern British Columbia, Canada, within the confines of Yoho National Park. At an elevation of 1,256 m (4,121 ft), it is 27 km (17 mi) west of Lake Louise along the Trans-Canada Highway which provides the only road access to the town.
Pikaia gracilens is an extinct, primitive chordate animal known from the Middle Cambrian Burgess Shale of British Columbia. Sixteen specimens are known from the Greater Phyllopod bed, where they comprised 0.03% of the community. It resembled the lancelet and perhaps swam much like an eel.
Waptia fieldensis is an extinct species of arthropod from the Middle Cambrian Burgess Shale Lagerstätte of Canada. It grew to a length of about 8 cm (3 in) and resembled modern shrimp in both morphology and habit. It had a large bivalved carapace and a segmented body terminating into a pair of tail flaps. It was an active swimmer, feeding on organic particles it gathers from the seafloor substrate. It is also one of the oldest animals with direct evidence of brood care.
Mount Burgess, 2,599 m (8,527 ft), is a mountain in Yoho National Park and is part of the Canadian Rockies. It is located in the southwest buttress of Burgess Pass in the Emerald River and Kicking Horse River Valleys.
Choia is a genus of extinct demosponge ranging from the Cambrian until the Lower Ordovician periods. Fossils of Choia have been found in the Burgess Shale in British Columbia; the Maotianshan shales of China; the Wheeler Shale in Utah; and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott.
Chancelloria is a genus of early animals known from the Middle Cambrian Burgess Shale, the Comley limestone, the Wheeler Shale, the Bright Angel Shale and elsewhere. It is named after Chancellor Peak. It was first described in 1920 by Charles Doolittle Walcott, who regarded them as one of the most primitive groups of sponges. This appears unlikely, and it is currently placed in the enigmatic group Coeloscleritophora. 178 specimens of Chancelloria are known from the Greater Phyllopod bed, where they comprise 0.34% of the community.
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 60,000 specimens in a series of field trips up from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to discover new species, and statistical analysis suggests discoveries will continue for the foreseeable future. Stephen Jay Gould's book Wonderful Life describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested.
The Stephen Formation is a geologic formation exposed in the Canadian Rockies of British Columbia. Its rocks were formed during the middle Cambrian. It is famous for the exceptional preservation of soft-bodied fossils: the Burgess Shale biota. The formation overlies the Cathedral escarpment, a submarine cliff; consequently it is divided into two quite separate parts, the 'thin' sequence deposited in the shallower waters atop the escarpment, and the 'thick' sequence deposited in the deeper waters beyond the cliff. Because the 'thick' Stephen Formation represents a distinct lithofacies, some authors suggest it warrants its own name, and dub it the Burgess Shale Formation. The stratigraphy of the Thin Stephen Formation has not been subject to extensive study, so except where explicitly mentioned this article applies mainly to the Thick Stephen Formation.
The Burgess Shale, a series of fossil beds in the Canadian Rockies, was first noticed in 1886 by Richard McConnell of the Geological Survey of Canada (GSC). His and subsequent finds, all from the Mount Stephen area, came to the attention of palaeontologist Charles Doolittle Walcott, who in 1907 found time to reconnoitre the area. He opened a quarry in 1910 and in a series of field trips brought back 65,000 specimens, which he identified as Middle Cambrian in age. Due to the quantity of fossils and the pressures of his other duties at the Smithsonian Institution, Walcott was only able to publish a series of "preliminary" papers, in which he classified the fossils within taxa that were already established. In a series of visits beginning in 1924, Harvard University professor Percy Raymond collected further fossils from Walcott's quarry and higher up on Fossil Ridge, where slightly different fossils were preserved.
Hazelia is a genus of spicular Cambrian demosponge known from the Burgess Shale, the Marjum formation of Utah, and possibly Chengjiang. It was described by Charles Walcott in 1920.
Takakkawia is a genus of sponge in the order Protomonaxonida and the family Takakkawiidae. It is known from the Middle Cambrian Burgess Shale that reached around 4 cm in height. Its structure comprises four columns of multi-rayed, organic spicules that align to form flanges. The spicules form blade-like structures, ornamented with concentric rings.
The Phyllopod bed, designated by USNM locality number 35k, is the most famous fossil-bearing member of the Burgess shale fossil Lagerstätte. It was quarried by Charles Walcott from 1911–1917, and was the source of 95% of the fossils he collected during this time; tens of thousands of soft-bodied fossils representing over 150 genera have been recovered from the Phyllopod bed alone.
Mackenzia is an elongated bag-like animal known from the Middle Cambrian Burgess Shale. It attached directly to hard surfaces, such as brachiopod shells. 14 specimens of Mackenzia are known from the Greater Phyllopod bed, where they comprise <0.1% of the community. Mackenzia was originally described by Charles Walcott in 1911 as a holothurian echinoderm. Later, Mackenzia is thought to be a cnidarian and appears most similar to modern sea anemones.
Louisella is a genus of worm known from the Middle Cambrian Burgess Shale. It was originally described by Charles Walcott in 1911 as a holothurian echinoderm, and represents a senior synonym of Miskoia, which was originally described as an annelid. 48 specimens of Louisella are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community. It has been stated to have palaeoscolecid-like sclerites, though this is not in fact the case.
The Mount Stephen trilobite beds are a series of fossil strata on Mount Stephen, British Columbia that contain exceptionally preserved fossil material. Part of the same stratigraphic unit as the Burgess Shale, many non-mineralized parts are preserved; in addition, a high density of trilobite fossils is present.
The Walcott Quarry is the most famous quarry of the Middle Cambrian Burgess Shale, located in the Canadian Rockies of British Columbia, bearing the Phyllopod beds. This lies at the base of the Walcott Quarry member, on a ridge between Wapta Mountain and Mount Field, and three other quarries – the Raymond, UE and EZ – lie above it. The quarry's proximity to the Cathedral escarpment led to the preservation of spectacular fossils.
Portalia mira is a fossil from the Burgess Shale that was originally described by Charles Walcott as a holothurian echinoderm. A later interpretation by F. Madsen in 1957 posits P. mira as a sponge.
Kutoringates are early rhynchonelliform brachiopods.
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