Reudemannoceratidae Temporal range: Middle Ordovician | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Cephalopoda |
Subclass: | Nautiloidea |
Order: | † Discosorida |
Family: | † Reudemannoceratidae Flower, 1940 |
The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center.
The Reudemannoceratidae first appeared at the beginning of the Middle Ordovician, North American Whiterock Stage, (since renamed to Dapingian by the International Commission on Stratigraphy), and are restricted to the lower part of that series: the middle Ordovician. Their origin is unknown. The siphuncles in early members contain features in the early growth stages reminiscent of the siphuncular bulbs found the archaic Plectronoceratae of the Late Cambrian. [1] but so far no unambiguous Lower Ordovician intermediaries have been found.
Reudemannoceratids are characterized by having short septal necks in the juvenile portion of the siphuncle, toward the apex of the shell, which later in life grow folded back along the back side of the septa in the characteristic fashion of the Discosorida. The connecting ring is bowed out into the camerae and is divided into the characteristic zones, the vinculum, granular zone, conchiolinous (or chitinous) zone offset by amorphous bands, and the layered bullette attached to the previous septal neck. [1] [2]
The Reucenammanoceratidae contain three genera. [2] They are Reudemannoceras, Franklinoceras, and Madiganella. Reudemannoceras and Franklinoceras, named by Flower in 1940 and 1957, are found the Champlain Valley in eastern North America. Madiganella, named by Teichert and Glenister in 1952, comes from central Australia.
Reudemannoceras is described [1] as having a somewhat compressed endogastric shell, such that the width is greater than the height, with the venter slightly flattened. Sutures are closely spaced and slope forward from venter to dorsum with the obliquity increasing as growth progressed. The early part of the shell is essentially straight, but afterward expands and is notably curved. The early segments in the siphuncle [2] are bulb shaped and the septal necks are short. In the later part of the shell she segments are expanded and septal necks become recumbent. The connecting rings are thick and have the zoning characteristic of the earlier Discosorids with well-developed bullettes.
Franklinoceras [2] is similar to Reudemannocereras, except that the shell is compressed and the sutures are straight.
Madiganella has a large, slender, straight or nearly straight shell [2] with a siphuncle composed of broad, expanded segments and short, strongly recurved necks. Growth lines indicate a shallow hyponomic sinus.
Reudemannoceras gave rise to the Cyrtogomphoceratidae through Ulrichoceras [2] as a result of an evolutionary ventral shift of the siphuncle and the development of large, inflated bullettes. Ulrichoceras is the probable ancestor of the Westonoceratidae as well. Reudemannoceras is also thought to have given rise [2] to Madiganella by an evolutionary straightening of the shell which by the subsequent development of T-shaped constricted apertures gave rise to the Mandaloceratidae. Franklinoceras seems to be an offshoot of Reudemanoceras with no progeny.
Reudemannoceratids probably lived on or near the sea floor. Endogastric Reudemannoceras and Franklinoceras are likely to have carried their shells high and to the back, aperture facing downward, as they probed the sea bed for prey. The more advanced and slightly later Madiganella may have been a fair swimmer, as indicated by the hyponomic sinus, and may have been an active stalker with a horizontal orientation.
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.
Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.
The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic).
The Lowoceratidae is a small family of discosorids, early nautiloid cephalopods, from the Middle Silurian in which the characteristic bullette is found only in early growth stages. Lowoceratids were first found in Southampton Island in the Canadian arctic.
Winnipegoceras is an extinct nautiloid genus from the Ordovician belonging to the Order Discosorida.
Ulrichoceras is recognised as the basal cyrtogomphoceratid genus, which is the source for the rest of the Cyrtogomphoceratidae as well as for the Westonoceratidae. The Cyrtogomphoceratidae are endogastric, the Westonoceratidae, exogastric.
Westonoceras is an extinct nautiloid genus from the Discosorida that lived during the Middle and Late Ordovician that has been found in North America, Greenland, and Northern Europe. It is the type genus for the Westonoceratidae
The Phragmoceratidae is a family of extinct nautiloid cephalopods from the Order Discosorida that lived during the latter part of the Silurian.
Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.
Mandaloceratidae is a family in the nautiloid cephalopod order Discosorida, from the Middle and Upper(?) Silurian characterized by short, essentially straight shells referred to as breviconic, typically with a faintly exogastric shape produced by the profile of the body chamber.
Landeroceras is a genus of straight shelled cyrtogomphoceratid from the Middle Ordovician Big Horn dolomite of Wyoming.
The Cyrtogomphoceratidae are a family in the cephalopod order Discosorida that comprises genera commonly with compressed, endogastrically curved shells. Siphuncles lie close to the ventral side, segments are broadly inflated, connecting rings thick and apically expanded thick bullettes. Chambers are short, separated by shallow, dish shaped septa. Apertures are generally simple.
Strandoceras is a strongly curved cyrtogomphoceratid with a laterally compressed cross section from the middle and upper Ordovician of Northern Europe; Estonia, Norway, the U.K.
Teichertoceras is a discosorid genus in the family Westonoceratidae characterized by an endogastric curvature to the early portion of the phragmocone.
Hectoceras is a genus in the nautiloid cephalopod order Discosorida from the Upper Ordovician of Australia (Tasmania), known from a few isolated siphuncle specimens.
Apocrinoceratidae constitutes a family of Middle Ordovician nautiloid cephalopods characterized by straight or slightly curved, transversely ribbed shells having siphuncles composed of expanded segments, short recurved septal necks, and thick connecting rings. Derivation is from the Protocycloceratidae, a family of ellesmerocerids, which differ in having straight or concave siphuncle segments, but are otherwise similar in form.
Cyrtogomphoceras is a genus of nautiloid cephalopods, recognized by its large breviconic shell with a notable endogastric curvature. The shell is fusiform in profile, reaching maximum width at or near the base of body chamber, which narrows toward the aperture. The siphuncle is large and slightly removed from the ventral side, that with the concave longitudinal profile. Siphuncle segments are short, as are chambers; septal necks recurved, connecting rings thick, bullettes at the apical end of the rings swollen. Cameral deposits are lacking.
Kiaeroceras is a slender, nearly straight shelled, cyrogomphoceratid (Nautiloidea-Discosorida) from the Upper Ordovician of northern Europe. The cross section of the shell is compressed, height greater than width. The body chamber is slightly contracted so as to narrow toward the aperture, which in some is slightly flared. The venter, narrowly rounded. The siphuncle is close to the venter, septal necks short, connecting rings thick, bullettes prominent.
Discosoridae comprise a family of endogastric discosorids, with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian.