Westonoceratidae | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Cephalopoda |
Subclass: | Nautiloidea |
Order: | † Discosorida |
Family: | † Westonoceratidae Flower & Teichert 1957 |
Westonoceratidae are exogastric, mostly compressed, Discosorida of moderate size from the Middle Ordovician to the Lower Silurian.
The siphuncle is typically close to the convexly curved outer margin of the phragmocone – the chambered part of the shell – taken to be ventral but may be more central in some genera. Connecting rings are thin to moderately thick with inflated bullettes grasping the previous septal foremina. Parietal deposits within the siphuncle from endocones in advanced genera and cameral deposits are found in some.
The Westonoceratidae are derived from the Cyrtogomphoceratidae through the cyrtogomphoceratid, Ulrichoceras . In turn, the Westonoceratidae gave rise to a small group of discosorids, the Lowoceratidae, which form an intermediary step before the Discosoridae.
Two principal lineages have been identified, both stemming from the genus Teichertoceras , a derivative of Ulrichoceras. One goes simply from Westonoceras to Winnipegoceras . The other goes from Faberoceras to Clarkesvillia to Glyptodendron . Faberoceras may also have given rise to Hecatoceras and Reedsoceras and is the likely source for the Lowoceratidae. Two other genera are included, but are separately derived from Ulrichoceras. These are Sinclairoceras and Simardoceras .
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.
The Tarphycerida were the first of the coiled cephalopods, found in marine sediments from the Lower Ordovician to the Middle Devonian. Some, such as Aphetoceras and Estonioceras, are loosely coiled and gyroconic; others, such as Campbelloceras, Tarphyceras, and Trocholites, are tightly coiled, but evolute with all whorls showing. The body chamber of tarphycerids is typically long and tubular, as much as half the length of the containing whorl in most, greater than in the Silurian Ophidioceratidae. The Tarphycerida evolved from the elongated, compressed, exogastric Bassleroceratidae, probably Bassleroceras, around the end of the Gasconadian through forms like Aphetoceras. Close coiling developed rather quickly, and both gyroconic and evolute forms are found in the early middle Canadian.
The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center.
The Lowoceratidae is a small family of discosorids, early nautiloid cephalopods, from the Middle Silurian in which the characteristic bullette is found only in early growth stages. Lowoceratids were first found in Southampton Island in the Canadian arctic.
Winnipegoceras is an extinct nautiloid genus from the Ordovician belonging to the Order Discosorida.
Ulrichoceras is recognised as the basal cyrtogomphoceratid genus, which is the source for the rest of the Cyrtogomphoceratidae as well as for the Westonoceratidae. The Cyrtogomphoceratidae are endogastric, the Westonoceratidae, exogastric.
Westonoceras is an extinct nautiloid genus from the Discosorida that lived during the Middle and Late Ordovician that has been found in North America, Greenland, and Northern Europe. It is the type genus for the Westonoceratidae
The Phragmoceratidae is a family of extinct nautiloid cephalopods from the Order Discosorida that lived during the latter part of the Silurian.
Mandaloceratidae is a family in the nautiloid cephalopod order Discosorida, from the Middle and Upper(?) Silurian characterized by short, essentially straight shells referred to as breviconic, typically with a faintly exogastric shape produced by the profile of the body chamber.
The Cyrtogomphoceratidae are a family in the cephalopod order Discosorida that comprises genera commonly with compressed, endogastrically curved shells. Siphuncles lie close to the ventral side, segments are broadly inflated, connecting rings thick and apically expanded thick bullettes. Chambers are short, separated by shallow, dish shaped septa. Apertures are generally simple.
Clarkesvillia is a genus of Upper Ordovician westonocerids the differs from Faberoceras in having a more flattened venter and from the later Glyptodendron in having a more ventrally located siphuncle.
Teichertoceras is a discosorid genus in the family Westonoceratidae characterized by an endogastric curvature to the early portion of the phragmocone.
Hectoceras is a genus in the nautiloid cephalopod order Discosorida from the Upper Ordovician of Australia (Tasmania), known from a few isolated siphuncle specimens.
Reedsoceras is a genus of large discosorids (Nautiloidea) in the family Westonoceratidae from the middle and upper Ordovician of North America.
Sinclairoceras is a nautiloid cephalopod in the discosorid family Westonoceratidae from the Middle Ordovician Simard Limestone of Quebec. The shell is laterally compressed, upwardly curved, and short; an exogastric brevicone with the venter narrowly and dorsum broadly rounded. The dorsum, at the upper surface, is only slightly concave longitudinally. The venter, at the lower surface, is more strongly convex longitudinally, drawing away from the dorsum and forming a hump at the anterior end of the phragmocone where Sinclairoceras reaches its greatest width.
Apocrinoceratidae constitutes a family of Middle Ordovician nautiloid cephalopods characterized by straight or slightly curved, transversely ribbed shells having siphuncles composed of expanded segments, short recurved septal necks, and thick connecting rings. Derivation is from the Protocycloceratidae, a family of ellesmerocerids, which differ in having straight or concave siphuncle segments, but are otherwise similar in form.
Cyrtogomphoceras is a genus of nautiloid cephalopods, recognized by its large breviconic shell with a notable endogastric curvature. The shell is fusiform in profile, reaching maximum width at or near the base of body chamber, which narrows toward the aperture. The siphuncle is large and slightly removed from the ventral side, that with the concave longitudinal profile. Siphuncle segments are short, as are chambers; septal necks recurved, connecting rings thick, bullettes at the apical end of the rings swollen. Cameral deposits are lacking.
Discosoridae comprise a family of endogastric discosorids,, with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian.
Kayoceras is a genus of middle Silurian nautiloid cephalopods, known from Iowa and Illinois, belonging to the Discosoridae. Related genera include Discosorus, Endodiscosorus, and Stokesoceras. Kayoceras has a short, breviconic shell, like Discosorus, but with a more central siphuncle. Derivation is probably from Discosorus, through an evolutionary shifting of the siphuncle position, which has its origin in the Lowoceratidae, possibly through Tuyloceras.