Winnipegoceras

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Winnipegoceras
Temporal range: M-U Ordovician
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Nautiloidea
Order: Discosorida
Family: Westonoceratidae
Genus: Winnipegoceras
Foerste (1922)

Winnipegoceras is an extinct nautiloid genus from the Ordovician belonging to the Order Discosorida.

The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.2 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.

Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.

Contents

Phylogeny

Winnipegoceras is included in the discosorid family Westonoceratidae and is derived from Westonoceras . Westonoceratidae have mostly medium to large, compressed, exogastric shells with moderately sized siphuncles composed of short folded back septal necks and generally thin connecting rings with swollen bullettes and which contain internal linings. (Flower & Teichert 1957; Teichert 1964)

Westonoceratidae are exogastric, mostly compressed, Discosorida of moderate size from the Middle Ordovician to the Lower Silurian.

Westonoceras is an extinct nautiloid genus from the Discosorida that lived during the Middle and Late Ordovician that has been found in North America, Greenland, and Northern Europe. It is the type genus for the Westonoceratidae

Siphuncle Strand of tissue passing longitudinally through the shell of a cephalopod mollusk

The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and Spirula. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the walls dividing the chambers.

Description

Winnipegoceras, named by Foerste (1922) for fossils found near Winnipeg in Manitoba, Canada, is known by it large, slender, compressed, strongly curved exogastric shells; long slender body chamber and siphuncle that is slightly removed from the venter which is the outer, longitudinally convex margin. Siphuncle segments are strongly narrowed at the septal foremina, bullettes swollen, parietal and other internal deposits rarely preserved. (ibid)

Winnipeg Provincial capital city in Manitoba, Canada

Winnipeg is the capital and largest city of the province of Manitoba in Canada. Centred on the confluence of the Red and Assiniboine rivers, it is near the longitudinal centre of North America, approximately 110 kilometres (70 mi) north of the Canada–United States border.

Manitoba Province of Canada

Manitoba is a province at the longitudinal centre of Canada. It is often considered one of the three prairie provinces and is Canada's fifth-most populous province with its estimated 1.3 million people. Manitoba covers 649,950 square kilometres (250,900 sq mi) with a widely varied landscape, stretching from the northern oceanic coastline to the southern border with the United States. The province is bordered by the provinces of Ontario to the east and Saskatchewan to the west, the territories of Nunavut to the north, and Northwest Territories to the northwest, and the U.S. states of North Dakota and Minnesota to the south.

Parietal callus

A parietal callus is a feature of the shell anatomy of some groups of snails, i.e. gastropods. It is a thickened calcareous deposit which may be present on the parietal wall of the aperture of the adult shell. The parietal wall is the margin of the aperture and part of the wall of the body whorl that is closest to the columella. The callus is often smooth and glossy, but can also be decorated with raised ribs or wrinkles.

Related Research Articles

Plectronocerida order of molluscs

Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.

Discosorus, a genus of the Discosorida and member of the family Discosoridae. Not to be confused with Discoceras. Discosorus consists of rapidly expanding endogastric brevicones, mostly known from isolated siphuncles composed of broadly expanded segments that increase rapidly in size. Septal necks are recumbent, connecting rings thin, bullettes small. The siphuncle interior is occupied by endocones produced by layered annular deposits expanded back toward the apex, leaving a central tube running down the middle. The short phragmocone is poorly known.

The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center.

The Lowoceratidae is a small family of discosorids, early nautiloid cephalopods, from the Middle Silurian in which the characteristic bullette is found only in early growth stages. Lowoceratids were first found in Southampton Island in the Canadian arctic.

Homaloceras is an extinct nautiloid cephalopod from the Middle Devonian with a strongly curved shell, included in the nautilid family Centroceratidae.

Ulrichoceras is recognised as the basal cyrtogomphoceratid genus, which is the source for the rest of the Cyrtogomphoceratidae as well as for the Westonoceratidae. The Cyrtogomphoceratidae are endogastric, the Westonoceratidae, exogastric.

Phragmoceratidae family of molluscs

The Phragmoceratidae is a family of extinct nautiloid cephalopods from the Order Discosorida that lived during the latter part of the Silurian.

Landeroceras is a genus of straight shelled cyrtogomphoceratid from the Middle Ordovician Big Horn dolomite of Wyoming.

The Cyrtogomphoceratidae are a family in the cephalopod order Discosorida that comprises genera commonly with compressed, endogastrically curved shells. Siphuncles lie close to the ventral side, segments are broadly inflated, connecting rings thick and apically expanded thick bullettes. Chambers are short, separated by shallow, dish shaped septa. Apertures are generally simple.

Teichertoceras is a discosorid genus in the family Westonoceratidae characterized by an endogastric curvature to the early portion of the phragmocone.

Hectoceras is a genus in the nautiloid cephalopod order Discosorida from the Upper Ordovician of Australia (Tasmania), known from a few isolated siphuncle specimens.

Reedsoceras is a genus of large discosorids (Nautiloidea) in the family Westonoceratidae from the middle and upper Ordovician of North America.

Simardoceras is a genus in the discosorid family Westonoceratidae from the Middle Ordovician of Quebec.

Sinclairoceras is a nautiloid cephalopod in the discosorid family Westonoceratidae from the Middle Ordovician Simard Limestone of Quebec. The shell is laterally compressed, upwardly curved, and short; an exogastric brevicone with the venter narrowly and dorsum broadly rounded. The dorsum, at the upper surface, is only slightly concave longitudinally. The venter, at the lower surface, is more strongly convex longitudinally, drawing away from the dorsum and forming a hump at the anterior end of the phragmocone where Sinclairoceras reaches its greatest width.

Cyrtogomphoceras is a genus of nautiloid cephalopods, recognized by its large breviconic shell with a notable endogastric curvature. The shell is fusiform in profile, reaching maximum width at or near the base of body chamber, which narrows toward the aperture. The siphuncle is large and slightly removed from the ventral side, that with the concave longitudinal profile. Siphuncle segments are short, as are chambers; septal necks recurved, connecting rings thick, bullettes at the apical end of the rings swollen. Cameral deposits are lacking.

Kiaeroceras is a slender, nearly stright shelled, cyrogomphoceratid (Nautiloidea-Discosorida) from the Upper Ordovician of northern Europe. The cross section of the shell is compressed, height greater than width. The body chamber is slightly contracted so as to narrow toward the aperture, which in some is slightly flared. The venter, narrowly rounded. The siphuncle is close to the venter, septal necks short, connecting rings thick, bullettes prominent.

Discosoridae comprise a family of endogastric discosorids,, with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian.

References

The Treatise on Invertebrate Paleontology published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by more than 300 paleontologists, and covering every phylum, class, order, family, and genus of fossil and extant invertebrate animals. The prehistoric invertebrates are described as to their taxonomy, morphology, paleoecology, stratigraphic and paleogeographic range. However, genera with no fossil record whatsoever have just a very brief listing.

See also