Mandaloceratidae

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Mandaloceratidae
Temporal range: M -?U Silurian
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Nautiloidea
Order: Discosorida
Family: Mandaloceratidae
Flower, 1957
Genera

See text

Mandaloceratidae is a family in the nautiloid cephalopod order Discosorida, from the Middle and Upper(?) Silurian characterized by short, essentially straight shells referred to as breviconic, typically with a faintly exogastric shape produced by the profile of the body chamber.

Contents

Apertures vary from round to T-shaped, with a long narrow, ventral, hyponomic sinus for the maneuvering funnel. Siphuncles are commonly central or subcentral with generally broad, expanded, segments and usually thin connecting rings.

The narrow hyponomic sinus, which is an opening for the water-jet funnel by which the animal could move, much as with modern squid and octopods, points toward the lower or ventral side. The cross bar of the "T" or the round part of the aperture through which the animal could extend itself is located closer to the back or dorsal part of the shell.

Phylogeny

The Mandaloceratidae are derived from the Reudemannoceratidae through Madiganella. The earliest mandeloceratid is Pseudogomphoceras which gave rise to Ovocerina. Ovocerina gave rise to Cinctoceras, Umbeloceras, Vesperoceras, and Mandaloceras. It also gave rise to the Mesoceratidae representing Mesoceras.

Genera

Mandaloceras is the type genus of the Mandaloceratidae characterized by straight or somewhat curved shells with a slightly depressed cross section (width greater than height) and a constricted T-shaped aperture. The siphncle is subcentral and increases rapidly in width during the life of the animal. Segments are broad with thin connecting rings.

Cinctoceras is a large mandaloceratid with a breviconic shell that tends to be dorsally flattened, making its cross section slightly depressed; a constricted T-shape aperture and siphuncle segments that are short and broad.

Ovocerina is a small breviconic mandaloceratid with an exogastric aspect produced by the more convexly rounded lower (ventral) side and a rounded or transversely extended aperture that approaches the T-shaped aperture of Mandaloceras. The siphuncle is displaced slightly, either ventrally or dorsally from the center. Segments, composed of thin connecting rings, are broadly expanded.

Pseudogomphoceras is a large, atypically longiconic mandeloceratid that reflects its reudemannoceratid ancestry. The subcentral siphuncle contains vesicular deposits and a central tube. Otherwise it is like Ovoverina.

Umbelloceras is a small breviconic mandaloceratid with a cross section ranging from slightly depressed to slightly compressed (width > height to width < height) and a T-shaped aperture in which the two lateral branches curve toward the venter.

Vesperoceras is a medium size, generally straight, breviconic mandaloceratid with a broad annular expansion to the anterior part of the phragmocone followed by a constriction at the base of the body chamber. The siphuncle is located midway between the center and venter. Segments are moderately to strongly expanded. The aperture may be open or constricted.

Mandaloceras was named by Hyatt in Zittel 1900. The rest were named by Rousseau Flower, Ovocerina in 1947, Cinctoceras, Pseudogomphoceras, Umbelloceras, and Vesperoceras in Flower and Teichert, 1957. With the exception of Mandaloceras, which is also found in North America, the Mandaloceratidae are restricted to the Silurian of Europe.

Related Research Articles

<span class="mw-page-title-main">Ascocerida</span> Extinct order of molluscs

The Ascocerida are comparatively small, bizarre Orthoceratoids known only from Ordovician and Silurian sediments in Europe and North America, uniquely characterized by a deciduous conch consisting of a longiconic juvenile portion and an inflated breviconic adult portion that separate sometime in maturity.

Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.

<span class="mw-page-title-main">Slender Oncoceratidae</span> Group of nautiloids

Slender Oncoceratidae are those in the family Oncoceratidae, which have slender, commonly curved, shells. Some like Oocerina are gently curved, almost straight, and with only slight expansion. Others like Dunleithoceras are strongly curved with a more notable rate of expansion. Inclusion in this somewhat arbitrary category is based on illustrations in the Treatise Part K, 1964.

The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center.

The Lowoceratidae is a small family of discosorids, early nautiloid cephalopods, from the Middle Silurian in which the characteristic bullette is found only in early growth stages. Lowoceratids were first found in Southampton Island in the Canadian arctic.

Turnoceras is a genus of Devonian cephalopods belonging to the oncocerid family Nothoceratidae. Its shell is broadly expanding and exogastrically curved such that the flattened dorsum is on longitudinally concave side. Aperture unconstricted with no hyponomic sinus for the water jet funnel. siphuncle along the outer, ventral, side, with radial, plate-like actinosiphonate deposits occupying the interior.

Brevicoceras is an extinct nautiloid genus from the order Oncocerida with wide distribution in the Middle Devonian in Eastern North America, Russia and Morocco. Nautiloids form a broad group of shelled cephalopods that were once diverse and numerous but are now represented by only a handful of species in two genera.

Ulrichoceras is recognised as the basal cyrtogomphoceratid genus, which is the source for the rest of the Cyrtogomphoceratidae as well as for the Westonoceratidae. The Cyrtogomphoceratidae are endogastric, the Westonoceratidae, exogastric.

Acleistoceras is a genus of the oncocerid, nautiloid family Acleistoceratidae that lived in the shallow seas that covered much of North America during the Devonian; living from 409—383.7 mya, existing for approximately 25.3 million years.

<span class="mw-page-title-main">Oncoceratidae</span> Extinct family of nautiloids

Oncoceratidae is a family of nauatiloid cephalopods in the order Oncocerida established by Hyatt, 1884, that range from the Middle Ordovician to the Upper Silurian.

Hemiphragmoceratidae is a family of endogastrically brevconic oncocerids characterized by elaborately visored apertures in which the hyponomic sinus in mature specimens is on a spout-like process and there may be lateral and dorsal salients.. Shells are compressed with the apical portion curved and the anterior straight. Siphucles are nummuloideal with expanded spheroidal segments and continuously actinosiphonate interiors.

<span class="mw-page-title-main">Phragmoceratidae</span> Extinct family of molluscs

The Phragmoceratidae is a family of extinct nautiloid cephalopods from the Order Discosorida that lived during the latter part of the Silurian.

The Cyrtogomphoceratidae are a family in the cephalopod order Discosorida that comprises genera commonly with compressed, endogastrically curved shells. Siphuncles lie close to the ventral side, segments are broadly inflated, connecting rings thick and apically expanded thick bullettes. Chambers are short, separated by shallow, dish shaped septa. Apertures are generally simple.

Pentameroceras is a straight to slightly exogastric breviconic oncocerid from the middle Silurian of North America and Europe belonging to the Trimeroceratidae.

<i>Ophioceras</i> Extinct genus of nautiloids

Ophioceras is a genus of closely coiled tarphycerid nautiloid cephalopods, the sole representatives of the family Ophidioceratidae, characterized by an evolute shell with narrow, subrounded, annulated whorls and a subcentral siphuncle composed of thin connecting rings that show no evidence of layering. The mature body chamber is strongly divergent and is the longest proportionally of any tarphycerid. The aperture has a deep hyponomic sinus and ocular sinuses, and so resembles some lituitids.

Poterioceratidae is a family of nautiloid cephalopods included in the Oncocerida that lived during the period from the Early Devonian to the Early Carboniferous (Mississippian). Members of the Poterioceratidae are distinguished by a subcircular to compressed exogastric shell that has no hyponomic sinus and a central to subcentral siphuncle composed of subquadrate to nummuloidal segments in which the septal necks are more strongly curved on the upper, or dorsal side. This is opposite from the Karoceratidae in which siphuncle segments are inflated ventrally but straight dorsally. Some poterioceratid genera have actinosiphonate structures or annular deposits within the siphuncle. In others it is empty.

Simardoceras is a genus in the discosorid family Westonoceratidae from the Middle Ordovician of Quebec.

Sinclairoceras is a nautiloid cephalopod in the discosorid family Westonoceratidae from the Middle Ordovician Simard Limestone of Quebec. The shell is laterally compressed, upwardly curved, and short; an exogastric brevicone with the venter narrowly and dorsum broadly rounded. The dorsum, at the upper surface, is only slightly concave longitudinally. The venter, at the lower surface, is more strongly convex longitudinally, drawing away from the dorsum and forming a hump at the anterior end of the phragmocone where Sinclairoceras reaches its greatest width.

Cyrtogomphoceras is a genus of nautiloid cephalopods, recognized by its large breviconic shell with a notable endogastric curvature. The shell is fusiform in profile, reaching maximum width at or near the base of body chamber, which narrows toward the aperture. The siphuncle is large and slightly removed from the ventral side, that with the concave longitudinal profile. Siphuncle segments are short, as are chambers; septal necks recurved, connecting rings thick, bullettes at the apical end of the rings swollen. Cameral deposits are lacking.

Discosoridae comprise a family of endogastric discosorids,, with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian.

References