River kingfisher | |
---|---|
A male common kingfisher (Alcedo atthis). | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Coraciiformes |
Family: | Alcedinidae |
Subfamily: | Alcedininae Rafinesque, 1815 |
Genera | |
Phylogeny of the Alcedininae | |||||||||||||||||||||
| |||||||||||||||||||||
Cladogram based on Andersen et al. (2017) [1] |
The river kingfishers or pygmy kingfishers, subfamily Alcedininae, are one of the three subfamilies of kingfishers. The river kingfishers are widespread through Africa and east and south Asia as far as Australia, with one species, the common kingfisher (Alcedo atthis) also appearing in Europe and northern Asia. This group includes many kingfishers that actually dive for fish. The origin of the subfamily is thought to have been in Asia.
These are brightly plumaged, compact birds with short tails, large heads, and long bills. They feed on insects or fish, and lay white eggs in a self-excavated burrow. Both adults incubate the eggs and feed the chicks.
A molecular phylogenetic study of the river kingfishers published in 2007 found that the genera as then defined did not form monophyletic groups. [2] The species were subsequently rearranged into four monophyletic genera. A clade containing four species were placed in the resurrected genus Corythornis and five species (little kingfisher, azure kingfisher, Bismarck kingfisher, silvery kingfisher and indigo-banded kingfisher) were moved from Alcedo to Ceyx. [3]
All except one of the kingfishers in the reconstituted Ceyx have three rather than the usual four toes. The exception is the Sulawesi dwarf kingfisher which retains a vestigial fourth toe. [2] [4]
The subfamily includes 35 species divided into four genera. The African dwarf kingfisher is sometimes placed in the monotypic genus Myioceyx, and sometimes with the pygmy kingfishers in Ispidina . Molecular analysis suggests that the Madagascar pygmy kingfisher is most closely related to the malachite kingfisher. [5]
All kingfishers are short-tailed large-headed compact birds with long pointed bills. Like other Coraciiformes, they are brightly coloured. Alcedo species typically have metallic blue upperparts and head, and orange or white underparts. The sexes may be identical, as with Bismarck kingfisher, but most species show some sexual dimorphism, ranging from a different bill colour as with common kingfisher to a completely different appearance. The male blue-banded kingfisher has white underparts with a blue breast band, whereas the female has orange underparts. [6]
The small kingfishers that make up the rest of the family have blue or orange upperparts and white or buff underparts, and show little sexual variation. [6] Across the family, the bill colour is linked to diet. The insectivorous species have red bills, and the fish-eaters have black bills. [7]
When perched, kingfishers sit quite upright, and the flight is fast and direct. The call is typically a simple high-pitched squeak, often given in flight. [8]
Most alcedinids are found in the warm climates of Africa and southern and southeast Asia. Three species reach Australia, but only the common kingfisher is found across most of Europe and temperate Asia. No members of this family are found in the Americas, although the American green kingfishers are believed to have derived from alcedinid stock. The origin of the family is thought to have been in southern Asia, which still has the most species. [9]
The Ceyx and Ispidina species are mainly birds of wet rainforest or other woodland, and are not necessarily associated with water. The Alcedo kingfishers are usually closely associated with fresh water, often in open habitats although some are primarily forest birds. [8]
River kingfishers are monogamous and territorial. The pair excavates a burrow in an earth bank and lays two or more white eggs onto the bare surface. Both parents incubate the eggs and feed the chicks. [8] Egg laying is staggered at one-day intervals so that if food is short only the older larger nestlings get fed. The chicks are naked, blind and helpless when they hatch, and stand on their heels, unlike any adult bird. [10]
The small Ceyx and Ispidina species feed mainly on insects and spiders, but also take tadpoles, frogs and mayfly nymphs from puddles. They will flycatch, and their red bills are flattened to assist in the capture of insects. The Alcedo kingfishers are typically fish-eaters with black bills, but will also take aquatic invertebrates, spiders and lizards. A few species are mainly insectivorous and have red bills. Typically fish are caught by diving into the water from a perch, although the kingfisher might hover briefly. [7]
Kingfishers are a family, the Alcedinidae, of small to medium-sized, brightly coloured birds in the order Coraciiformes. They have a cosmopolitan distribution, with most species living in the tropical regions of Africa, Asia, and Oceania, but also can be seen in Europe. They can be found in deep forests near calm ponds and small rivers. The family contains 116 species and is divided into three subfamilies and 19 genera. All kingfishers have large heads, long, sharp, pointed bills, short legs, and stubby tails. Most species have bright plumage with only small differences between the sexes. Most species are tropical in distribution, and a slight majority are found only in forests.
The common kingfisher, also known as the Eurasian kingfisher and river kingfisher, is a small kingfisher with seven subspecies recognized within its wide distribution across Eurasia and North Africa. It is resident in much of its range, but migrates from areas where rivers freeze in winter.
The malachite kingfisher is a river kingfisher which is widely distributed in Africa south of the Sahara. It is largely resident except for seasonal climate-related movements.
The half-collared kingfisher is a kingfisher in the subfamily Alcedininae that is found in southern and eastern Africa. It feeds almost exclusively on fish and frequents streams, rivers and larger bodies of water with dense shoreline vegetation.
The American pygmy kingfisher is a species of "water kingfisher" in subfamily Cerylinae of family Alcedinidae. It is found in the American tropics from southern Mexico south through Central America into every mainland South American country except Chile and Uruguay. It also occurs on Trinidad.
The African pygmy kingfisher is a small insectivorous kingfisher found in the Afrotropics, mostly in woodland habitats.
Pelargopsis is a genus of tree kingfishers that are resident in tropical south Asia from India and Sri Lanka to Indonesia.
Alcedo is a genus of birds in the kingfisher subfamily Alcedininae. The genus was introduced by Carl Linnaeus in 1758 in the 10th edition of his Systema Naturae. The type species is the common kingfisher. Alcedo is the Latin for "kingfisher".
The little kingfisher is a species of kingfisher in the subfamily Alcedininae.
The black-backed dwarf kingfisher, also known as the three-toed kingfisher, is a pocket-sized bird in the family Alcedinidae. It was formerly considered as conspecific with the rufous-backed dwarf kingfisher and together the species complex was known by the English name "oriental dwarf kingfisher".
The African dwarf kingfisher is a species of kingfisher in the Alcedininae subfamily.
Ceyx is an Old World genus of river kingfishers. These kingfishers are found from South East Asia to the Solomon Islands.
Ispidina is a genus of small insectivorous African river kingfishers.
The tree kingfishers, also called wood kingfishers or Halcyoninae, are the most numerous of the three subfamilies of birds in the kingfisher family, with around 70 species divided into 12 genera, including several species of kookaburras. The subfamily appears to have arisen in Indochina and Maritime Southeast Asia and then spread to many areas around the world. Tree kingfishers are widespread through Asia and Australasia, but also appear in Africa and the islands of the Pacific and Indian Oceans, using a range of habitats from tropical rainforest to open woodlands.
Corythornis is a genus of small African river kingfishers.
The dimorphic dwarf kingfisher is a species of bird in the family Alcedinidae that is endemic to the central and southern Philippines. Its natural habitat is subtropical or tropical moist lowland forests.
The Buru dwarf kingfisher, is a species of bird in the family Alcedinidae that is endemic to Buru Island in Indonesia. Its natural habitat is subtropical or tropical moist lowland forests.
The Papuan dwarf kingfisher, is a species of bird in the family Alcedinidae that is endemic to the western Papuan islands, New Guinea, Aru Islands and the D'Entrecasteaux Archipelago. Its natural habitat is subtropical or tropical moist lowland forests.
The North Solomons dwarf kingfisher, is a species of bird in the family Alcedinidae that is endemic to the west and central Solomon Islands. Its natural habitat is subtropical or tropical moist lowland forests.
The Malaita dwarf kingfisher, is a subspecies of bird in the family Alcedinidae that is endemic to Malaita Island. Its natural habitat is subtropical or tropical moist lowland forests.