| Rotaciurca Temporal range: Late Silurian, | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Hemichordata |
| Class: | Pterobranchia |
| Order: | Cephalodiscida |
| Family: | † Rotaciurcidae Briggs & Mongiardino Koch, 2023 |
| Genus: | † Rotaciurca Briggs & Mongiardino Koch, 2023 |
| Species: | †R. superbus |
| Binomial name | |
| †Rotaciurca superbus Briggs & Mongiardino Koch, 2023 | |
Rotaciurca superbus is a Silurian cephalodiscid pterobranch, known from the Bertie Group of Ontario. [1] [2]
Rotaciurca is a pseudocolonial (containing multiple clustered individuals which are not connected) pterobranch, unique for being a cephalodiscid rather than a graptolite. The cluster consists of a ring of radiating tubes, likely containing the living individuals, attached to a conical structure interpreted as a float, giving the original fossil the nickname of "Ezekiel's wheel". [2] The whole organism is roughly 4 cm (1.6 in) long, seemingly with two distinct rows of tubes. Its pterobranch affinity is confirmed by lineations within the tubes, which are traces of fuselli, and the irregular arrangement of these group it with the cephalodiscids. Rotaciurca has a number of similarities to Yuknessia , another enigmatic fossil recently described as a pterobranch, including rigid and similarly sized tubes, although those of Yuknessia are much finer and have more tightly packed fuselli. [1]
Rotaciurca has been interpreted as a planktonic filter-feeder, with the conical float providing buoyancy like that of a Portuguese man o' war. It is unique in this aspect, as all other known cephalodiscids are benthic and sessile, while the only other planktonic pterobranchs are graptolites, which shows the water column was colonised several times by hemichordates. [1]
Rotaciurca is known from ten specimens, with the holotype nearly complete. Unusually for a planktonic animal, it has only been found in one locality, that being the Bertie Group, however since it was likely soft-bodied, the exceptional preservation of this site may have been necessary for Rotaciurca to fossilise. [1]
Rotaciurca is named in honour of Samuel J. Ciurca, Jr., who donated thousands of eurypterid fossils to the Yale Peabody Museum, alongside the Latin word rota, meaning "wheel". Its specific name, superbus, from the Latin word meaning "beautiful", acknowledges that Ciurca labelled the holotype fossil "the most beautiful fossil ever found". [1]
A chordate is a deuterostomal bilaterian animal belonging to the phylum Chordata. All chordates possess, at some point during their larval or adult stages, five distinctive physical characteristics (synapomorphies) that distinguish them from other taxa. These five synapomorphies are a notochord, a hollow dorsal nerve cord, an endostyle or thyroid, pharyngeal slits, and a post-anal tail.
Hemichordata is a phylum which consists of triploblastic, eucoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and include two main classes: Enteropneusta, and Pterobranchia. A third class, Planctosphaeroidea, is known only from the larva of a single species, Planctosphaera pelagica. The class Graptolithina, formerly considered extinct, is now placed within the pterobranchs, represented by a single living genus Rhabdopleura.
Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian through the Lower Carboniferous (Mississippian). A possible early graptolite, Chaunograptus, is known from the Middle Cambrian. Recent analyses have favored the idea that the living pterobranch Rhabdopleura represents an extant graptolite which diverged from the rest of the group in the Cambrian.
Chitinozoa are a group of flask-shaped, organic walled marine microfossils produced by an as yet unknown organism. Common from the Ordovician to Devonian periods, the millimetre-scale organisms are abundant in almost all types of marine sediment across the globe. This wide distribution, and their rapid pace of evolution, makes them valuable biostratigraphic markers.
Pterygotus is a genus of giant predatory eurypterid, a group of extinct aquatic arthropods. Fossils of Pterygotus have been discovered in deposits ranging in age from Middle Silurian to Late Devonian, and have been referred to several different species. Fossils have been recovered from four continents; Australia, Europe, North America and South America, which indicates that Pterygotus might have had a nearly cosmopolitan (worldwide) distribution. The type species, P. anglicus, was described by Swiss naturalist Louis Agassiz in 1839, who gave it the name Pterygotus, meaning "winged one". Agassiz mistakenly believed the remains were of a giant fish; he would only realize the mistake five years later in 1844.
Pterobranchia, members of which are often called pterobranchs, is a class of small worm-shaped animals. They belong to the Hemichordata, and live in secreted tubes on the ocean floor. Pterobranchia feed by filtering plankton out of the water with the help of cilia attached to tentacles. There are about 25 known living pterobranch species in three genera, which are Rhabdopleura, Cephalodiscus, and Atubaria. On the other hand, there are several hundred extinct genera, some of which date from the Cambrian Period.
Megalograptus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Megalograptus have been recovered in deposits of Katian age in North America. The genus contains five species: M. alveolatus, M. ohioensis, M. shideleri, M. welchi and M. williamsae, all based on fossil material found in the United States. Fossils unassigned to any particular species have also been found in Canada. The generic name translates to "great writing" and originates from the mistaken original belief that Megalograptus was a type of graptolite, often given names ending with -graptus.
Isotelus is a genus of asaphid trilobites from the Middle and Late Ordovician Period, fairly common in the northeastern United States, northwest Manitoba, southwestern Quebec and southeastern Ontario.
Judeasaurus is an extinct genus of small, aquatic varanoid lizard related to the mosasauroids. The only known specimen is from the Late Cretaceous of the Judean Hills around Israel and the West Bank, though its exact provenance is uncertain.
Slimonia is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Slimonia have been discovered in deposits of Silurian age in South America and Europe. Classified as part of the family Slimonidae alongside the related Salteropterus, the genus contains three valid species, S. acuminata from Lesmahagow, Scotland, S. boliviana from Cochabamba, Bolivia and S. dubia from the Pentland Hills of Scotland and one dubious species, S. stylops, from Herefordshire, England. The generic name is derived from and honors Robert Slimon, a fossil collector and surgeon from Lesmahagow.
Erettopterus is a genus of large predatory eurypterid, an extinct group of aquatic arthropods. Fossils of Erettopterus have been discovered in deposits ranging from Early Silurian to the Early Devonian, and have been referred to several different species. Fossils have been recovered from two continents; Europe and North America. The genus name is composed by the Ancient Greek words ἐρέττω (eréttō), which means "rower", and πτερόν (pterón), which means "wing", and therefore, "rower wing".
Rhinocarcinosoma is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Rhinocarcinosoma have been discovered in deposits ranging of Late Silurian age in the United States, Canada and Vietnam. The genus contains three species, the American R. cicerops and R. vaningeni and the Vietnamese R. dosonensis. The generic name is derived from the related genus Carcinosoma, and the Greek ῥινός, referring to the unusual shovel-shaped protrusion on the front of the carapace of Rhinocarcinosoma, its most distinctive feature.
Phoronids are a small phylum of marine animals that filter-feed with a lophophore, and build upright tubes of chitin to support and protect their soft bodies. They live in most of the oceans and seas, including the Arctic Ocean but excluding the Antarctic Ocean, and between the intertidal zone and about 400 meters down. Most adult phoronids are 2 cm long and about 1.5 mm wide, although the largest are 50 cm long.
Ambedus is an extinct genus of possible diadectid reptiliomorph. Fossils have been found from the Early Permian Dunkard Group of Monroe County, Ohio. The type species A. pusillus was named in 2004. The genus name comes from the Latin word ambedo meaning "to nibble", in reference to its herbivorous diet. The specific name pusillus means "tiny" in Latin.
Bellubrunnus is an extinct genus of rhamphorhynchid pterosaur from the Late Jurassic of southern Germany. It contains a single species, Bellubrunnus rothgaengeri. Bellubrunnus is distinguished from other rhamphorhynchids by its lack of long projections on the vertebrae of the tail, fewer teeth in the jaws, and wingtips that curve forward rather than sweep backward as in other pterosaurs.
Haootia quadriformis is an extinct animal belonging to the Ediacaran biota. Estimated to be about 560 million years old, H. quadriformis is identified as a cnidarian polyp, and represents the earliest known evidence for muscle tissue in an animal. Discovered in 2008 from Newfoundland in eastern Canada, it was formally described in 2014. It is the first Ediacaran organism discovered to show fossils of muscle fibres. Structural examination of the muscles and morphology indicate that the animal is a cnidarian, though, which class H. quadriformis belongs to was undetermined until a 2024 study found it to be a staurozoan.
Etacystis communis, colloquially known as the H-animal or aitch, was a soft-bodied invertebrate that lived in shallow tropical coastal waters of muddy estuaries during the Pennsylvanian geological period, about 300 million years ago. The classification is uncertain: the animal had a unique H-shaped body ranging from 2 to 11 cm long, and researchers have suggested a hemichordate or hydrozoan affinity. Examples of Etacystis have been found only in the Mazon Creek fossil beds of the Midwestern United States.
Necrogammarus salweyi is the binomial name applied to an arthropod fossil discovered in Herefordshire, England. The fossil represents a fragmentary section of the underside and an appendage of a pterygotid eurypterid, a group of large and predatory aquatic arthropods that lived from the late Silurian to the late Devonian. The Necrogammarus fossil is Late Silurian in age and its generic name means "dead lobster", deriving from Ancient Greek νεκρός and Latin gammarus ("lobster").
The Bertie Group or Bertie Limestone, also referred to as the Bertie Dolomite and the Bertie Formation, is an upper Silurian geologic group and Lagerstätte in southern Ontario, Canada, and western New York State, United States. Details of the type locality and of stratigraphic nomenclature for this unit as used by the U.S. Geological Survey are available on-line at the National Geologic Map Database. The formation comprises dolomites, limestones and shales and reaches a thickness of 495 feet (151 m) in the subsurface, while in outcrop the group can be 60 feet (18 m) thick.
Luchibang is an extinct genus of istiodactylid pterosaur discovered in Inner Mongolia, China. The type and only species is Luchibang xingzhe. The genus was initially established on the basis of a supposed nearly complete skeleton and skull, although later analysis of the specimen revealed that it represents a chimera, with the Luchibang holotype restricted to the end of the snout.