| Samadera harmandiana | |
|---|---|
Scientific classification  | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Angiosperms |
| Clade: | Eudicots |
| Clade: | Rosids |
| Order: | Sapindales |
| Family: | Simaroubaceae |
| Genus: | Samadera |
| Species: | S. harmandiana |
| Binomial name | |
| Samadera harmandiana | |
| Synonyms [1] | |
| |
Samadera harmandiana (syn Quassia harmandiana) is a freshwater mangrove shrub or small tree in the Simaroubaceae family. It is found in Cambodia, Laos and Vietnam. The wood provides firewood. Certain fish eat the poisonous fruit
This species grows as a shrub or small tree, some 5–8 m (16.4–26.2 ft) tall. [2]
The species was placed in the section Samadera by Noteboom in 1962, along with Quassia indica . [3] The plants of this section have simple leaves with roughly scattered concave glands, mostly on the under surface. The flowers are bisexual, occurring in axillary or terminal inflorescences, either pseudoumbels with peduncles or in racemes. There are 3-5 calyx lobes, these are imbricate in the bud, obtuse and have a concave gland in the centre. There are 3-5 contorted petals, much longer than the calyx, usually hairy on the underside. The disc is large, as high as it is broad, and gynophore-like. The style has a inconspicuous terminal stigma. The quite large fruits are compressed laterally, with a narrow, thinner, sharp-edged part in the apex half. Quassia harmandiana fruit are very large and dorsoventrally compressed. It flowers from March to May and fruits in July. [4] The fruit are poisonous, see below.
It was first published by Jean Baptiste Louis Pierre based on an earlier description by Jean Marie Antoine de Lanessan in Pl. Util. Col. Franç. on page 305 in 1886. [1]
The specific epithet name of Harmandiana is in honour of Julien Herbert Auguste Jules Harmand (1844-1915), who was a French clergyman and botanist (Mycology and Lichenology). [5]
The species was later placed in the section Samadera of the Simaroubaceae by Dutch botanist Hans Peter Nooteboom (1934–2022) in 1962 (published in 1963), as Quassia harmandiana, along with Quassia indica . Nooteboom had taken a very broad view of the genus Quassia L. and included therein various genera including, Hannoa Planch., Odyendyea (Pierre) Engl., Pierreodendron Engl., Samadera Gaertn., Simaba Aubl. and Simarouba Aubl. . [3]
In 2007, DNA sequencing, and phylogenetic analyses was carried out on members of the Simaroubaceae family. It found that genus Samadera was a sister to Clade V and that genus Quassia was also a sister to Clade V but they had separate lineages. This suggested the splitting up of genera Quassia again, with all Nooteboom's synonyms listed above being resurrected as independent genera. [6]
It was later returned to genus Samadera as Samadera harmandiana in 2007. [1] [7]
Krahs and kra: (Khmer) are names used in Cambodia. [2] Mak ngooan is a name from Laos. [8]
The species grows in the following countries: Cambodia, Laos and Vietnam. [1]
The plant grows in the mangrove and back-mangrove formations of the Mekong and Tonle Sap rivers and lake. [2]
In the vegetation communities alongside the Mekong in Kratie and Steung Treng Provinces, Cambodia, this taxa is moderately abundant in the dense diverse strand community (last to be flooded each year, first to drain). [4] It also occurs in mixed evergreen and deciduous forests in the area. It grows on soils derived from metamorphic sandstone bedrock, at of 25–30 m (82–98 ft) altitudes above sea level.
In the Khone Falls section of the Mekong, in southern Laos, close to where the river becomes fully Cambodian, two species of carp-like fish, Leptobarbus hoevenii and Tor cf. tambra eat the fruits of this species. [8] [9] The flesh of these fish is rendered poisonous after eating this and other fruit. In Tor cf. tambra the eyes become red and scales become whiter if poison is present, and if recognised, then the fish can be made edible by discarding the head and guts, then dried in the sun or marinated.[ citation needed ]
The wood makes excellent firewood. [2] The plant contains bactericidal compounds.
Quassia is a plant genus in the family Simaroubaceae. Its size is disputed; some botanists treat it as consisting of only one species, Quassia amara from tropical South America, while others treat it in a wide circumscription as a pantropical genus containing up to 40 species of trees and shrubs.
Sindora siamensis is a species of tree in the subfamily Detarioideae of the family Fabaceae. It has an accepted infraspecific, the variety S. siamensis var. maritima (Pierre) K.Larsen & S.S.Larsen. See taxon box to the right below, and below for details on the variety maritima. The nominate species is found in many countries in tropical Asia. Like several other species in the genus Sindora, its wood is considered valuable; the least concern conservation status may reflect efforts to replant this species, but mortality rates are high. As well as the wood, the plant provides raw material for chemical products, food and drink, and domestic utensils.
Pierreodendron is a genus of plants in the family Simaroubaceae.
Elaeocarpus lanceifolius is a tree species in the family Elaeocarpaceae. It is found across tropical Asia from Thailand to Yunnan to Nepal to Karnataka, India. It is used for its wood, fruit, and nuts.
Ziziphus cambodiana is a deciduous thorny shrub, or vine, some 2–6 m tall, found growing in secondary undergrowth in Cambodia, Laos and Vietnam, and northern Thailand.
Brownlowia emarginata is a slightly climbing tree, a member of the family Malvaceae. It occurs in Vietnam, Cambodia, Laos and Thailand.
A small tree with tortuous twigs, Dillenia pentagyna is a member of the family Dilleniaceae, and is found from Sulawesi to South-Central China to India and Sri Lanka. Material from the tree has some minor uses.
Homalium brevidens is a shrub or tree species in the family Salicaceae, found in Laos and Cambodia.
Dendrolobium baccatum is a species of flowering plants in the Fabaceae family. A shrub, it occurs in Mainland Southeast Asia. People use it for food and fuel.
Erythroxylum cambodianum is a shrub in the family Erythroxylaceae. It grows in Vietnam, Laos, Cambodia and Thailand. The wood is used for pickets and as firewood.
Gluta cambodiana is a shrub/small tree in the family Anacardiaceae. It occurs in parts of Mainland Southeast Asia. Its wood is used for pickets and fuel.
Combretum trifoliatum is a vine of the family Combretaceae. It is found from Myanmar across Southeast Asia and Wallacea to New Guinea and Australia. It grows in wet places, including where it can be submerged for four months a year by floodwaters. It is unusual in retaining its photosynthesizing leaves when flooded. Parts of the plant are used in traditional medicine.
Memecylon caeruleum is a shrub or tree species in the Melastomataceae family. It is found from New Guinea, west through Southeast Asia to Tibet, Zhōngguó/China. It has become an invasive weed in the Seychelles. It has some local use for wood and food.
Pantadenia adenanthera is a shrub in the Euphorbiaceae family. It is found in parts of Southeast Asia. The species is used for its wood and edible fruit.
Stixis obtusifolia is a shrub or liana in the Resedaceae family. It is found in parts of Southeast Asia. The wood is used as fuel, the leaves as a tea.
Xanthophyllum lanceatum is a tree in the Polygalaceae family. It grows across Southeast Asia from Sumatera to Bangladesh. The leaves are used as a hops-substitute in beer making and the wood as fuel. Fish in the Mekong regularly eat the fruit, flowers and leaves.
Ampelocissus martini is a species of climber or shrub in the Vitaceae family. Some sources use the spelling Ampelocissus martinii. It is native to an area of Mainland Southeast Asia. The fruit are eaten by people and by several species of Pangasiidae shark catfish of the Mekong river.
Memecylon lilacinum is a tree species in the Melastomataceae family. It is usually an understorey species in closed forests. It is native to an area of tropical Asia, from Jawa to Philippines to Vietnam and the Andaman Islands and Myanmar. It is a food plant for the macaque Macaca facsicularis and a bee in the Megachilidae family.
Simaba is a genus of flowering plants belonging to the family Simaroubaceae.
Samadera is a genus of trees belonging to the family Simaroubaceae, in the order Sapindales. The genus is native to eastern Africa (Tanzania), across to parts of tropical Asia and parts of eastern Australia.