Steelhead in 1924 illustration using the original taxonomic name, Salmo gairdneri
The freshwater form of the steelhead is the rainbow trout (Oncorhynchus mykiss). The difference between these forms of the species is that steelhead migrate to the ocean and return to freshwater tributaries to spawn, whereas non-anadromous rainbow trout do not leave freshwater. Steelhead are also larger and less colorful than rainbow trout. Steelhead can weigh up to 55lb (25kg) and reach 45in (110cm) in length. They can live up to 11 years and spawn multiple times.[4] The body of the steelhead trout is silvery and streamlined with a rounder head. This silver color and round head is what gives the steelhead its name.[4] There are black dots that are more concentrated on the back of the fish and become sparser closer to the lateral line of the fish. Steelhead also develop a pink horizontal stripe. When steelhead return to freshwater to spawn, their color begins to more closely resemble that of a normal rainbow trout. The longer a steelhead spends in freshwater, the darker their green body colorations and horizontal pink stripe become. When a steelhead exits the river and returns to the ocean again, it regains its silvery appearance while in saltwater. The fish develop a larger and stronger tail relative to a normal rainbow trout to help aid in swimming in ocean currents and swimming up strong river currents.
A number of distinct population segments of steelhead are endangered or threatened across the United States, mostly caused by the blocking of waterways by the construction of dams. Human interaction has had considerable consequences on reducing the population of steelhead trout.[5]
Characteristics
Steelhead currently support or have historically supported fisheries across Northern California, and thus they hold "economic as well as cultural value" (Moyle, Israel, & Purdy, 2008). Contrary to popular belief, the California Steelhead is a coastal variation of the same species of fish as the Rainbow Trout, Oncorhynchus mykiss; "rainbow trout are the 'landlocked' version, and remain in freshwater throughout their life" (King County, 2016). An ocean-going variety, the Steelhead, or Oncorhynchus mykiss irideus has developed unique adaptive characteristics that distinguish this fish from its common counterpart and have allowed a wider distribution of this species across the Pacific Northwest, including the coastal regions of California. In fact, "coastal rainbow trout is the most widely-distributed native trout form" (CA.gov). Among other necessary biological utilities fulfilled by this species, Steelhead Trout "are also strong indicators of the condition of California's streams; large self-sustaining populations of native salmon and trout are found where streams are in reasonably good condition" (Moyle, Israel, & Purdy, 2008). A chrome underside and olive-grey back reminiscent to that of a steel beam are attributed as the main defining physical characteristic for which the Steelhead is distinguished and earns its namesake.
Steelhead trout drift in an Oregon stream.
The Steelhead is an ocean-going predatory fish with a typical lifespan of four to six years. Like the common rainbow trout, Steelhead predominantly feast on zooplankton when they are young and then transition to eating fish, some rodents, mollusks, and crustaceans. When these fish enter the open ocean, however, their diet typically consists of squid, crustaceans, and small fish including anchovies, herring, and sardines, though the capacity at which this dietary intake occurs is highly contingent upon the geographical region where Steelhead choose to migrate (Salmon Fishing Now). "Rainbow trout and steelhead represent two divergent ecotypes that are genetically identical but separated by life history strategies" (Heath 2001); the unique biological diversity of the Steelhead Trout from that of the Rainbow Trout is determined through a capacity to migrate to the open ocean. Homing behavior is a unique adaptation of Steelhead Trout that allows the fish to migrate to and from freshwater and saltwater bodies on multiple occasions. Through the process of imprinting, Steelhead Trout deposit a chemical indicator "that is specific to their natal stream" (NRC 1996).[6][7]
Homing behavior
Perhaps the most interesting and perplexing characteristic of the Steelhead Trout is its homing behavior and the capacity of this species to accurately detect its primary spawning grounds despite venturing to the open ocean and returning to the same freshwater source on numerous occasions throughout their lives. By recording the concentrations of solutes and the chemical profile of a stream through the biological characteristic of a chemical indicator, steelhead "use the position of the sun and magnetic north to navigate towards spawning grounds " after they have fed in the open ocean for 1–4 whole years (Fulton). Even after traveling hundreds of miles, this chemical imprint allows steelhead to identify their precise hatching grounds upon returning to freshwater. According to "A review of the characteristics, habitat requirements, and ecology of the Anadromous Steelhead Trout" by Fulton, "homing creates genetically different subpopulations that differ by basin, stream, and even location within a stream" (Altukhov 2000). According to a study conducted by Heath & Pollard, "significant genetic variation on the individual, tributary, and watershed level occur between Steelhead Trout populations" as a result of this chemical reliance for breeding practices (Heath, 2001). The author concludes that as a result of such biologically accurate homing characteristics, the Steelhead Trout is "uniquely adapted to its environment"; fish populations "are reasonably isolated from fish that spawn in different areas," and therefore "sub-populations may differ by the timing of runs, size of fish, and even behavior patterns" (Heath, 2001). This also influences genetic relatedness between Steelhead Trout populations; interestingly, the author found that the genetic similarity of regional Steelhead trout populations "is correlated with distance" (Heath, 2001).[8]
During spawning, the fish will lay their eggs on gravel bottoms of freshwater tributaries. The female digs out a hole called a redd. The preferred depth for steelhead spawning is 6 to 14 inches (150 to 360mm).[9] She then lays the eggs, and a male fertilizes them. The females will then cover the eggs with the gravel. Depending on the size of the female she may lay up to 9,000 eggs. The female then buries the eggs in a foot of gravel. The eggs remain in the gravel until they hatch. Unlike salmon (which can spawn only once) steelhead may return to the ocean, and come back upstream several times for spawning season.
Great Lakes steelhead
Steelhead found in the Great Lakes are the freshwater variation of steelhead found in saltwater. Similar to native populations of steelhead on the west coast, Great Lakes steelhead start in tributaries to larger bodies of water, in this case the Great Lakes, where they travel to, to grow to large proportions and reach sexual maturity before returning to the tributaries they started in. Great Lakes steelhead grow to as large as 20 pounds and 36–40 inches but normally average 4–10 pounds and between 20–30 inches.[10] They share very similar characteristics to an ocean-run steelhead as far as their long, slender build and silvery coloration with spots running along their back that very rarely progress beyond their lateral line. As the fish spend more time in the tributaries they darken in coloration while developing a bright red stripe down their midline. While in the lakes they feed on alewives, smelt, and lake herring amongst other things.
The first introduction of steelhead into the Great Lakes dates back to 1876[11] where smolts were taken from wild populations from the McCloud River in California and placed into the Au Sable River. "Wild" populations of these fish started to become established shortly after their introduction into the lakes. By the late 80s populations could be found in notable rivers like the Muskegon and Pere Marquette. Sometime after their first introduction into Michigan tributaries other states, such as New York, Ohio, and Pennsylvania, started stocking fish into their own streams.[citation needed] Due to the steelhead's large size and powerful demeanor, they've become very attractive amongst the angler community in the Great Lakes area.
Conservation status
There have been several conservation programs created since the near extinction of the steelhead in the 1940s. The reduction in population is mainly due to manmade obstructions within river systems. This is usually caused by dams blocking access, or humans changing the river landscape for recreation and access to water. It is estimated that only 500 steelhead trout return to the Southern California watersheds. However, in other areas of the United States, steelhead can be seen as pests because they are an invasive species to the environment.[5]
Southern California
While numbers are improving in areas of the Pacific Northwest where Steelhead Trout have typically thrived, the California Steelhead Trout is considered a threatened species, and according to Mark Capelli of the National Marine Fisheries Service, the Southern California steelhead are the most endangered of them all (California Trout). According to California Trout, Steelhead Trout populations began to decline in the "late-1940s due mainly to man-made landscape modifications" (California Trout). As a result of damming for flood control, steelhead migration routes from the coast to upstream spawning grounds were increasingly blocked over time (California Trout). Furthermore, largely a result of continued urbanization and to satisfy the needs of human populations in the area, river flows became modified for recreational purposes and a substantial diversion of water occurred over time (California Trout). As a result of widespread habitat loss and fragmentation, Southern California Steelhead populations were officially labeled as an endangered species in 1997 (California Trout). In some areas, a recent sighting of the Steelhead Trout has not occurred in years, and biologists fear the worst. In 2002, the endangered species label was extended from the Malibu Creek down to the border of San Diego and Mexico (California Trout).
Northern California
A threatened species since 2000, the Northern California steelhead represents a beacon of hope for continued rehabilitation initiatives and growing population numbers in the area. As of 2016, 48,892 Steelhead Trout were reportedly caught (NOAA). Such a population count suggests that this fascinating fish species continue to persevere in Northern California waters. However, to ensure the continued rehabilitation of steelhead populations in Northern California, a positive, directive plan of action for steelhead conservation must occur. According to NOAA, The State of California Fisheries Restoration Grant Program (FRGP) "invested over 250 million dollars and supported approximately 3,500 salmonid restoration projects" (NOAA). Investments in projects improving "fish passage, water quality, instream habitat restoration, watershed monitoring, and education and organizational support" all occupy conservation efforts to improve Northern California steelhead populations (NOAA). According to NOAA, "the percentage of floodplain and in-channel habitat that would need to be restored to detect a 25% increase in salmon and steelhead production was 20%" (NOAA). As it stands, "more than 20% of floodplain and in-channel habitat has been restored due to FRGP" (NOAA). Though these initiatives spell hope for the prospects of Steelhead Trout population rehabilitation, habitat degradation continues to occur at an alarming rate.
Oregon
Management Units and Population Segments for the MCR Steelhead.
Within the state of Oregon there are four different distinct steelhead population segments: Lower Columbia River steelhead, Middle Columbia River steelhead, Oregon Coast steelhead, and Upper Willamette River Steelhead[12] (NOAA). Of these four segments, three are listed as threatened under the Endangered Species Act. The Middle Columbia River (MCR) segment has remained the focus of recent recovery efforts. The MCR segment covers approximately 3,500 square miles of the Columbia Plateau in eastern Oregon and Washington.[13] Within eastern Oregon, recovery efforts focus on the two main tributaries to the Columbia River: The Deschutes River and the John Day River, although the Umatilla and Walla Walla are also included in recovery plans.[14] Primary threats to MCR steelhead populations include hydrosystem operations, habitat alteration, and effects of stray naturally spawning hatchery fish.[13] MCR steelhead were first listed as threatened on March 25, 1999, following the construction of Round Butte Dam at the confluence of the Deschutes and Metolius rivers.[13] The construction of the dam created reverse currents (currents flowing upstream) in the Metolius Arm of Lake Billy Chinook (a reservoir created by the formation of the dam) confusing anadromous smolts and preventing them from navigating seaward. The construction of Round Butte Dam impeded steelhead access to the Upper Deschutes, Crooked, and Metolius rivers, three major tributaries to the Deschutes within the MCR watershed.[13] In addition to Round Butte, several other dams restrict fish from their native spawning grounds within the Deschutes watershed including Pelton Dam, Bowman Dam, and Ochoco Dam. The first MCR Distinct Population Segment ESA Recovery Plan was released on November 30, 1999. Between 2010 and 2016 1,715 different entities have implemented 1,559 tributary restoration projects primarily focused on habitat restoration and fish passage improvement.[13] In a 2016 an action plan implementation progress report stated the population is "maintained" showing no significant upward or downward trend in the yearly number of returning adults in a experimental small tributary to the Deschutes River.[14] However, fish counts at Boneville Dam, on the Columbia River below the Deschutes confluence have shown a 10 year (2013–2022) average of 162,988 returning adults, and a four year (2019–2022) average of 97,352, suggesting a recent decline in rates of returning adults.[15]
↑ Behnke, Robert J. (2002). "Rainbow and Redband Trout". Trout and Salmon of North America. Tomelleri, Joseph R. (illustrator). New York: The Free Press. p.67. ISBN978-0-7432-2220-4.
↑ "Steelhead Trout". NOAA Fisheries. National Oceanic and Atmospheric Association. 17 February 2022.
↑ Behnke, Robert J. (2002). "Rainbow and Redband Trout". Trout and Salmon of North America. Tomelleri, Joseph R. (illustrator). New York: The Free Press. p.76. ISBN978-0-7432-2220-4.
Aaron Fulton. "A Review of the Characteristics, Habitat Requirements, and Ecology of the Anadromous Steelhead Trout (Oncorhynchus Mykiss) in the Skeena Basin," June 15, 2004, 16.
"Coastal Multispecies Plan Volume III: Northern California Steelhead." NOAA Fisheries, 2016. https://www.fisheries.noaa.gov/resource/document/final-coastal-multispecies-recovery-plan-california-coastal-chinook-salmon.
"Coastal Rainbow Trout/ Steelhead." California Department of Fish and Wildlife, October 29, 2016. https://wildlife.ca.gov/Conservation/Fishes/Coastal-Rainbow-Trout-Steelhead.
"Fish Hatcheries." In Fishing in Idaho. Idaho Department of Fish and Game, n.d. https://idfg.idaho.gov/visit/hatchery.
Peter B. Moyle, Joshua A. Israel, and Sabra E. Purdy. "Salmon, Steelhead, and Trout in California: Status of an Emblematic Fauna." California Trout, 2008, 220.
"Rainbow Trout and Steelhead." National Wildlife Federation, n.d. https://www.nwf.org/Educational-Resources/Wildlife-Guide/Fish/Rainbow-Trout-Steelhead.
"Southern Steelhead: A Story of Recovery." California Trout, February 7, 2018. https://caltrout.org/news/southern-steelhead-story-recovery.
"Steelhead Trout." In NOAA Fisheries. NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION, n.d. https://www.fisheries.noaa.gov/species/steelhead-trout.
"Steelhead Trout Identification." King County, November 10, 2016. https://www.kingcounty.gov/services/environment/animals-and-plants/salmon-and-trout/identification/steelhead.aspx.
V. Kuhnlein, Harriet, and Murray M. Humphries. "Rainbow Trout (Steelhead Trout)." In Traditional Animal Foods of Indigenous Peoples of Northern North America. Centre for Indigenous Peoples' Nutrition and Environment, n.d. http://traditionalanimalfoods.org/fish/freshwater/page.aspx?id=6151.
Salmon is the common name for several commercially important species of euryhaline ray-finned fish from the genera Salmo and Oncorhynchus of the family Salmonidae, native to tributaries of the North Atlantic (Salmo) and North Pacific (Oncorhynchus) basins. Other closely related fish in the same family include trout, char, grayling, whitefish, lenok and taimen, all coldwater fish of the subarctic and cooler temperate regions with some sporadic endorheic populations in Central Asia.
Trout is a generic common name for numerous species of carnivorous freshwater ray-finned fishes belonging to the genera Oncorhynchus, Salmo and Salvelinus, all of which are members of the subfamily Salmoninae in the family Salmonidae. The word trout is also used for some similar-shaped but non-salmonid fish, such as the spotted seatrout/speckled trout.
Fish migration is mass relocation by fish from one area or body of water to another. Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few metres to thousands of kilometres. Such migrations are usually done for better feeding or to reproduce, but in other cases the reasons are unclear.
The Napa River is a river approximately 55 miles (89 km) long in the U.S. state of California. It drains a famous wine-growing region called the Napa Valley, in the mountains north of the San Francisco Bay. Milliken Creek and Mt. Veeder watersheds are a few of its many tributaries. The river mouth is at Vallejo, where the intertidal zone of fresh and salt waters flow into the Carquinez Strait and the San Pablo Bay.
The rainbow trout is a species of trout native to cold-water tributaries of the Pacific Ocean in North America and Asia. The steelhead is an anadromous (sea-run) form of the coastal rainbow trout(O. m. irideus) or Columbia River redband trout (O. m. gairdneri) that usually returns to freshwater to spawn after living two to three years in the ocean. Freshwater forms that have been introduced into the Great Lakes and migrate into tributaries to spawn are also called steelhead.
Oncorhynchus is a genus of ray-finned fish in the subfamily Salmoninae of the family Salmonidae, native to coldwater tributaries of the North Pacific basin. The genus contains twelve extant species, namely six species of Pacific salmon and six species of Pacific trout, all of which are migratory mid-level predatory fish that display natal homing and semelparity.
Puget Sound salmon recovery is a collective effort of federal, state and local authorities and non-profit coalitions of universities, scientists, business and industry aimed at restoring Pacific salmon and anadromous forms of Pacific trout (Oncorhynchus) within the Puget Sound region. The Puget Sound lies within the native range of the Pacific Salmon (Oncorhynchus) and two sea-run forms of Pacific trout, the coastal rainbow trout or steelhead and coastal cutthroat trout. Populations of Oncorhynchus have seen significant declines since the middle of the 19th century due to over fishing, habitat loss, pollution and disease. Salmon species residing in or migrating through the Puget Sound to spawning streams include Chum, Coho, Chinook, Sockeye, and Pink salmon. Pacific salmon require freshwater rivers for spawning and most major tributaries of the Puget Sound have salmon, steelhead and cutthroat trout spawning runs.
Redband trout are a group of three recognized subspecies of rainbow trout. They occur in three distinct regions in Pacific basin tributaries and endorheic basins in the western United States. The three subspecies are the Columbia River redband trout, the McCloud River redband trout and the Great Basin redband trout.
Alameda Creek is a large perennial stream in the San Francisco Bay Area. The creek runs for 45 miles (72 km) from a lake northeast of Packard Ridge to the eastern shore of San Francisco Bay by way of Niles Canyon and a flood control channel. Along its course, Alameda Creek provides wildlife habitat, water supply, a conduit for flood waters, opportunities for recreation, and a host of aesthetic and environmental values. The creek and three major reservoirs in the watershed are used as water supply by the San Francisco Public Utilities Commission, Alameda County Water District and Zone 7 Water Agency. Within the watershed can be found some of the highest peaks and tallest waterfall in the East Bay, over a dozen regional parks, and notable natural landmarks such as the cascades at Little Yosemite and the wildflower-strewn grasslands and oak savannahs of the Sunol Regional Wilderness.
San Francisquito Creek is a creek that flows into southwest San Francisco Bay in California, United States. Historically it was called the Arroyo de San Francisco by Juan Bautista de Anza in 1776. San Francisquito Creek courses through the towns of Portola Valley and Woodside, as well as the cities of Menlo Park, Palo Alto, and East Palo Alto. The creek and its Los Trancos Creek tributary define the boundary between San Mateo and Santa Clara counties.
San Leandro Creek is a 21.7-mile-long (34.9 km) year-round natural stream in the hills above Oakland in Alameda County and Contra Costa County of the East Bay in northern California.
Arroyo Corte Madera del Presidio is a 4.1-mile-long (6.6 km) year-round stream in southern Marin County, California, United States. This watercourse is also known as Corte Madera Creek, although the actual stream of that name flows into San Francisco Bay further north at Point San Quentin. This watercourse has a catchment basin of about 8 square miles (21 km2) and drains the south-eastern slopes of Mount Tamalpais and much of the area in and around the town of Mill Valley; this stream discharges to Richardson Bay.
The coastal cutthroat trout, also known as the sea-run cutthroat trout, blue-back trout or harvest trout, is one of the four species of cutthroat trout found in Western North America. The coastal cutthroat trout occurs in four distinct forms. A semi-anadromous or sea-run form is the most well known. Freshwater forms occur in both large and small rivers and streams and lake environments. The native range of the coastal cutthroat trout extends south from the southern coastline of the Kenai Peninsula in Alaska to the Eel River in Northern California. Coastal cutthroat trout are resident in tributary streams and rivers of the Pacific basin and are rarely found more than 100 miles (160 km) from the ocean.
Marsh Creek is a stream in east Contra Costa County, California in Northern California which rises on the eastern side of Mount Diablo and flows 30 miles (48 km) to the Sacramento–San Joaquin River Delta at Oakley, California, near Big Break Regional Shoreline. The creek flows through Marsh Creek State Park (California), where water is impounded to form Marsh Creek Reservoir, then through the city of Brentwood, California.
Searsville Dam is a masonry dam in San Mateo County, California, that was completed in 1892, one year after the founding of Stanford University, and impounds Corte Madera Creek to form a reservoir known as Searsville Reservoir or Searsville Lake. Searsville Dam is located in the Jasper Ridge Biological Preserve and is owned and operated by Stanford University. Neighboring cities include Woodside and Portola Valley, California.
Corte Madera Creek is a 7.3-mile-long (11.7 km) creek that flows north-northwest to Searsville Dam and then joins with Bear Creek to form San Francisquito Creek in California.
The Kamchatkan rainbow trout is a subspecies of the rainbow trout, which is a fish in the family Salmonidae. It is native to Russian Far East. Its main range is on the Kamchatka Peninsula, and it has also been recorded from the Commander Islands east of Kamchatka, and sporadically in the Sea of Okhotsk, as far south as the mouth of the Amur River.
The Kern River rainbow trout is a localized subspecies of the rainbow trout, a variety of fish in the family Salmonidae. It is found in a short section of the main stem of the Kern River and several tributaries in the southern Sierra Nevada in California. The Kern River rainbow trout is a "Species of Special Concern" in the state of California due to habitat loss and hybridization with other native and non-native trout in their range.
Old Mill Creek is a 2.3-mile-long (3.7 km) year-round stream in southern Marin County, California, United States. Old Mill Creek is a tributary to Arroyo Corte Madera del Presidio, which, in turn, flows to Richardson Bay at Mill Valley's municipal Bayfront Park and 106 acres (0.43 km2) Bothin Marsh.
Little Arthur Creek, historically San Antonio Creek, is a 6.1-mile-long (9.8 km) east/southeastward-flowing stream originating just west of the crest of the Santa Cruz Mountains. It joins Uvas Creek about 4 miles (6.4 km) west of Gilroy, California, Santa Clara County, California. Uvas Creek is, in turn, tributary to the Pajaro River and thence to Monterey Bay and the Pacific Ocean.
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