The lacustrine and riverine morphs of brown trout are both potamodromous, meaning they are also migratory, though only between freshwater bodies. Lacustrine trout mainly inhabit large lakes with calm and stratified deep water, while riverine trout forms fluvial populations typically in large rivers but sometimes in shallower creeks and alpine streams, both still migrating upstream during reproductive seasons. Anadromous and potamodromous morphs coexisting in the same river appear genetically identical.[5] What determines whether they migrate to sea or not remains unknown.
Taxonomy
The scientific name of the brown trout is Salmo trutta. The specific epithet trutta derives from the Latintrutta, meaning, literally, "trout". Behnke (2007) relates that the brown trout was the first species of trout described in the 1758 edition of Systema Naturae by Swedish zoologist Carl Linnaeus. Systema Naturae established the system of binomial nomenclature for animals. Salmo trutta was used to describe anadromous or sea-run forms of brown trout. Linnaeus also described two other brown trout species in 1758. Salmo fario was used for riverine forms. Salmo lacustris was used for lake-dwelling forms.[6]
Brown trout have been widely introduced into suitable environments around the world, including North and South America, Australasia, Asia, and South and East Africa. Introduced brown trout have established self-sustaining, wild populations in many introduced countries.[8] The first introductions were in Australia in 1864 when 300 of 1500 brown trout eggs from the River Itchen survived a four-month voyage from Falmouth, Cornwall to Melbourne on the sailing ship Norfolk. By 1866, 171 young brown trout were surviving in a Plenty River hatchery in Tasmania. Thirty-eight young trout were released in the river, a tributary of the River Derwent in 1866. By 1868, the Plenty River hosted a self-sustaining population of brown trout which became a brood source for continued introduction of brown trout into Australian and New Zealand rivers.[9] Successful introductions into the Natal and Cape Provinces of South Africa took place in 1890 and 1892, respectively. By 1909, brown trout were established in the mountains of Kenya. The first introductions into the Himalayas in northern India took place in 1868, and by 1900, brown trout were established in Kashmir and Madras.[10] In the 1950s and 1960s, Edgar Albert de la Rue[fr], a French geologist, began the introduction of several species of salmonids on the remote Kerguelen Islands in the southern Indian Ocean. Of the seven species introduced, only brook trout, Salvelinus fontinalis, and brown trout survived to establish wild populations.[11]
Introduction to Americas
The first introductions in Canada occurred in 1883 in Newfoundland[12] and continued until 1933. The only Canadian regions without brown trout are Yukon and the Northwest Territories. Introductions into South America began in 1904 in Argentina. Brown trout are now established in Chile, Peru and the Falklands.[9] Sea-run forms of brown trout exceeding 20lb (9.1kg) are caught by local anglers on a regular basis.[citation needed]
U.S. range of brown trout
The first introductions into the U.S. started in 1883 when Fred Mather, a New York pisciculturist and angler, under the authority of the U.S. Fish Commissioner, Spencer Baird, obtained brown trout eggs from a Baron Lucius von Behr, president of the German Fishing Society[de]. The von Behr brown trout came from both mountain streams and large lakes in the Black Forest region of Baden-Württemberg.[7] The original shipment of "von Behr" brown trout eggs were handled by three hatcheries, one on Long Island, the Cold Spring Hatchery operated by Mather, one in Caledonia, New York, operated by pisciculturalist Seth Green, and other hatchery in Northville, Michigan. Additional shipments of "von Behr" brown trout eggs arrived in 1884. In 1885, brown trout eggs from Loch Leven, Scotland, arrived in New York. These "Loch Leven" brown trout were distributed to the same hatcheries. Over the next few years, additional eggs from Scotland, England, and Germany were shipped to U.S. hatcheries. Behnke (2007) believed all life forms of brown trout—anadromous, riverine, and lacustrine—were imported into the U.S. and intermingled genetically to create what he calls the American generic brown trout and a single subspecies the North European brown trout (S. t. trutta).[7]
In April 1884, the U.S. Fish Commission released 4900 brown trout fry into the Baldwin River, a tributary of the Pere Marquette River in Michigan. This was the first release of brown trout into U.S. waters. Between 1884 and 1890, brown trout were introduced into suitable habitats throughout the U.S.[7] By 1900, 38 states and two territories had received stocks of brown trout. Their adaptability resulted in most of these introductions establishing wild, self-sustaining populations.[9]
Conservation status
Infographic about the brown trout
The fish is not considered to be endangered, although some individual stocks are under various degrees of stress mainly through habitat degradation, overfishing, and artificial propagation leading to introgression. Increased frequency of excessively warm water temperatures in high summer causes a reduction in dissolved oxygen levels which can cause "summer kills" of local populations if temperatures remain high for sufficient duration and deeper/cooler or fast, turbulent more oxygenated water is not accessible to the fish. This phenomenon can be further exacerbated by eutrophication of rivers due to pollution—often from the use of agricultural fertilizers within the drainage basin.[citation needed]
Overfishing is a problem where anglers fail to identify and return mature female fish into the lake or stream. Each large female removed can result in thousands fewer eggs released back into the system when the remaining fish spawn.[citation needed]
In small streams, brown trout are important predators of macroinvertebrates, and declining brown trout populations in these specific areas affect the entire aquatic food web.[13]
Global climate change is also of concern. S. trutta morpha fario prefers well-oxygenated water in the temperature range of 60 to 65°F (16 to 18°C). S. trutta bones from an archaeological site in Italy, and ancient DNA extracted from some of these bones, indicate that both abundance and genetic diversity increased markedly during the colder Younger Dryas period, and fell during the warmer Bølling-Allerød event.[14]
Cover or structure is important to trout, and they are more likely to be found near submerged rocks and logs, undercut banks, and overhanging vegetation. Structure provides protection from predators, bright sunlight, and higher water temperatures. Access to deep water for protection in winter freezes, or fast water for protection from low oxygen levels in summer are also ideal. Trout are more often found in heavy and strong currents.[citation needed]
Defining characteristics include a slender body with a long, narrow head. The mouth is large, and on its roof, vomerine teeth are developed in a zig-zag pattern.[15][16] The caudal fin is deltaform without forking, unlike that of the related Atlantic salmon (Salmo salar).[15] Dark and red spots are often present on the sides, but do not extend to the tail.[16] Parr trout (juvenile) often have a red margin on their adipose fin, with dark blotches along their sides that also become inconspicuous with age.[16]
Freshwater brown trout range in colour from largely silver with relatively few spots and a white belly, to the more well-known brassy reddish-brown cast fading to creamy white on the fish's belly, with medium-sized spots surrounded by lighter halos. The more silver forms can be mistaken for rainbow trout. Regional variants include the so-called "Loch Leven" trout, distinguished by larger fins, a slimmer body, and heavy black spotting, but lacking red spots. The continental European strain features a lighter golden cast with some red spotting and fewer dark spots. Notably, both strains can show considerable individual variation from this general description. Early stocking efforts in the United States used fish taken from Scotland and Germany.
A 2.7-kg (6 lb), 60-cm (2 ft) sea trout, from Galway Bay in the west of Ireland bearing scars from a fishing netBrown trout in a creekBrown trout in Värmland, Sweden, after the first summerA young brown trout from the River Derwent in North East EnglandBrown trout from a western Wyoming creek
The brown trout is a medium-sized fish, growing to 20kg (44lb) or more and a length of about 100cm (39in) in some localities, although in many smaller rivers, a mature weight of 1.0kg (2.2lb) or less is common. S. t. lacustris reaches an average length of 40–80cm (16–31in) with a maximum length of 140cm (55in) and about 60 pounds (27kg).[citation needed]
On September 11, 2009, a 41.45-lb (18.80-kg) brown trout was caught by Tom Healy in the Manistee River system in Michigan, setting a new state record.[17] As of late December 2009, the fish captured by Healy was confirmed by both the International Game Fish Association and the Fresh Water Fishing Hall of Fame as the new all-tackle world record for the species. This fish, which supplanted the former world record from the Little Red River in Arkansas,[17] has in turn been exceeded by a 20.1-kilogram (44lb) specimen caught in the Ohau Canal in Twizel, New Zealand on 27 October 2020. The all-tackle length IGFA world record is a 97-centimetre (38in) fish caught in Milwaukee Harbor, Wisconsin on 16 December 2011.[18]
The spawning behaviour of brown trout is similar to that of the closely related Atlantic salmon. A typical female produces about 2,000 eggs per kg (900 eggs per lb) of body weight at spawning.
Brown trout can live 20 years, but as with the Atlantic salmon, a high proportion of males die after spawning, and probably fewer than 20% of anadromous female kelts recover from spawning [citation needed]. The migratory forms grow to significantly larger sizes for their age due to abundant forage fish in the waters where they spend most of their lives. Sea trout are more commonly female in less nutrient-rich rivers. Brown trout are active both by day and by night and are opportunistic feeders. While in freshwater, their diets frequently include invertebrates from the streambed, other fish, frogs, mice, birds, and insects flying near the water's surface. The high dietary reliance upon insect larvae, pupae, nymphs, and adults allows trout to be a favoured target for fly fishing. Sea trout are fished for especially at night using wet flies. Brown trout can be caught with lures such as spoons, spinners, jigs, plugs, plastic worm imitations, and live or dead baitfish.
Brown trout rarely form hybrids with other species; if they do, they are almost invariably infertile. One such example is the tiger trout, a hybrid with the brook trout.
Diet
Field studies have demonstrated that brown trout fed on several animal prey species, aquatic invertebrates being the most abundant prey items. However, brown trout also feed on other taxa such as terrestrial invertebrates (e.g. Hymenoptera) or other fish.[19] Moreover, in brown trout, as in many other fish species, a change in the diet composition normally occurs during the life of the fish,[20] and piscivorous behaviour is most frequent in large brown trout.[21] These shifts in the diet during fish lifecycle transitions may be accompanied by a marked reduction in intraspecific competition in the fish population, facilitating the partitioning of resources.[22][23]
First feeding of newly emerged fry is very important for brown trout survival in this phase of the lifecycle, and first feeding can occur even prior to emergence.[24][25] Fry start to feed before complete yolk absorption and the diet composition of newly emerged brown trout is composed of small prey such as chironomid larvae or baetid nymphs.[26]
The species has been widely introduced for sport fishing into North America, South America, Australia, New Zealand, and many other countries, including Bhutan, where they are the focus of a specialised fly fishery. The first planting in the United States occurred on April 11, 1884, into the Baldwin River, one mile east of Baldwin, Michigan.[27] Brown trout have had serious negative impacts on upland native fish species in some of the countries where they have been introduced, particularly Australia. In Chile, Australia, New Zealand and other locations in the southern hemisphere, brown trout compete with fish from the family Galaxiidae, which also have affinity for well-oxygenated, cold streams. Brown trout additionally are voracious predators of invertebrates and can carry microbial pathogens like Aeromonas salmonicida.[28] Genetic background is a very important factor when determining the success of trout populations, this information is vital to restore and enhance previous populations.[29] Because of the trout's importance as a food and game fish, it has been artificially propagated and stocked in many places in its range, and fully natural populations (uncontaminated by allopatricgenomes) probably exist only in isolated places, for example in Corsica or in high alpine valleys on the European mainland.[citation needed]
Farming of brown trout has included the production of infertile triploid fish by increasing the water temperature just after fertilisation of eggs, or more reliably, by a process known as pressure shocking. Triploids are favoured by anglers because they grow faster and larger than diploid trout. Proponents of stocking triploids argue, because they are infertile, they can be introduced into an environment that contains wild brown trout without the negative effects of cross-breeding. However, stocking triploids may damage wild stocks in other ways. Triploids certainly compete with diploid fish for food, space, and other resources. They could also be more aggressive than diploid fish and they may disturb spawning behaviour.[citation needed]
Angling
Frontis and title page from The Fly-fisher's Entomology, 1849, by Alfred Ronalds, showing a brown trout and a grayling
The brown trout has been a popular game fish of European anglers for centuries. It was first mentioned in angling literature as "fish with speckled skins" by Roman author Aelian (circa 200 AD) in On the Nature of Animals. This work is credited with describing the first instance of fly fishing for trout, the trout being the brown trout found in Macedonia.[30] The Treatyse of Fysshynge with an Angle (1496) by Dame Juliana Berners, O.S.B is considered a foundational work in the history of recreational fishing, especially fly fishing. One of the most prominent fish described in the work is the brown trout of English rivers and streams:
The trout, because he is a right dainty fish and also a right fervent biter, we shall speak of next. He is in season from March until Michaelmas. He is on clean gravel bottom and in a stream.
The renowned The Compleat Angler (1653) by Izaak Walton is replete with advice on "the trout":
The Trout is a fish highly valued, both in this and foreign nations. He may be justly said, as the old poet said of wine, and we English say of venison, to be a generous fish: a fish that is so like the buck, that he also has his seasons; for it is observed, that he comes in and goes out of season with the stag and buck. Gesner says, his name is of a German offspring; and says he is a fish that feeds clean and purely, in the swiftest streams, and on the hardest gravel; and that he may justly contend with all fresh water fish, as the Mullet may with all sea fish, for precedency and daintiness of taste; and that being in right season, the most dainty palates have allowed precedency to him.
Throughout the 17th, 18th, and 19th centuries, angling authors, mostly British, some French, and later American, writing about trout fishing were writing about fishing for brown trout. Once brown trout were introduced into the U.S. in the 1880s, they became a major subject of American angling literature. In 1889, Frederic M. Halford, a British angler, author published Dry-Fly Fishing in Theory and Practice, a seminal work codifying a half century of evolution of fly fishing with floating flies for brown trout. In the late 19th century, American angler and writer Theodore Gordon, often called the "Father of American Dry Fly Fishing", perfected dry-fly techniques for the newly arrived, but difficult-to-catch brown trout in Catskill rivers such as the Beaverkill and Neversink Rivers.[33] In the early 20th century, British angler and author G. E. M. Skues pioneered nymphing techniques for brown trout on English chalk streams. His Minor Tactics of the Chalk Stream (1910) began a revolution in fly fishing techniques for trout.[34] In 1917, Scottish author Hamish Stuart published the first comprehensive text, The Book of The Sea Trout, specifically addressing angling techniques for the anadromous forms of brown trout.[35]
Firehole river brown trout
Introductions of brown trout into the American West created new angling opportunities, none so successful from an angling perspective as was the introduction of browns into the upper Firehole River in Yellowstone National Park in 1890.[36] One of the earliest accounts of trout fishing in the park is from Mary Trowbridge Townsend's 1897 article in Outing Magazine "A Woman's Trout Fishing in Yellowstone Park" in which she talks about catching the von Behr trout in the river:
Long dashes down stream taxed my unsteady footing; the sharp click and whirr of the reel resounded in desperate efforts to hold him somewhat in check; another headlong dash, then a vicious bulldog shake of the head as he sawed back and forth across the rocks. Every wile inherited from generations of wily ancestors was tried until, in a moment of exhaustion, the net was slipped under him. Wading ashore with my prize, I had barely time to notice his size—a good four-pounder, and unusual markings, large yellow spots encircled by black, with great brilliancy of iridescent color—when back he flopped into the water and was gone. However, I took afterward several of the same variety, known in the Park as the Von Baer [sic] trout, and which I have since found to be the Salmo fario, the veritable trout of Izaak Walton.
Within the US, brown trout introductions have created self-sustaining fisheries throughout the country. Many are considered "world-class" such as in the Great Lakes and in several Arkansas tailwaters.[38] Outside the U.S. and outside its native range in Europe, introduced brown trout have created "world-class" fisheries in New Zealand,[39]Patagonia,[40] and the Falklands.[41]
Related Research Articles
Trout is a generic common name for numerous species of carnivorous freshwater ray-finned fishes belonging to the genera Oncorhynchus, Salmo and Salvelinus, all of which are members of the subfamily Salmoninae in the family Salmonidae. The word trout is also used for some similar-shaped but non-salmonid fish, such as the spotted seatrout/speckled trout.
Fly fishing is an angling technique that uses an ultralight-weight lure called an artificial fly, which typically mimics small invertebrates such as flying and aquatic insects to attract and catch fish. Because the mass of the fly lure is insufficient to overcome air resistance, it cannot be launched far using conventional gears and techniques, so specialized tackles are used instead and the casting techniques are significantly different from other forms of angling. It is also very common for the angler to wear waders, carry a hand net, and stand in the water when fishing.
The rainbow trout is a species of trout native to cold-water tributaries of the Pacific Ocean in Asia and North America. The steelhead is an anadromous (sea-run) form of the coastal rainbow trout(O. m. irideus) or Columbia River redband trout (O. m. gairdneri) that usually returns to freshwater to spawn after living two to three years in the ocean. Freshwater forms that have been introduced into the Great Lakes and migrate into tributaries to spawn are also called steelhead.
The Atlantic salmon is a species of ray-finned fish in the family Salmonidae. It is the third largest of the Salmonidae, behind Siberian taimen and Pacific Chinook salmon, growing up to a meter in length. Atlantic salmon are found in the northern Atlantic Ocean and in rivers that flow into it. Most populations are anadromous, hatching in streams and rivers but moving out to sea as they grow where they mature, after which the adults seasonally move upstream again to spawn.
The brook trout is a species of freshwater fish in the char genus Salvelinus of the salmon family Salmonidae. It is native to Eastern North America in the United States and Canada, but has been introduced elsewhere in North America, as well as to Iceland, Europe, and Asia. In parts of its range, it is also known as the eastern brook trout, speckled trout, brook charr, squaretail, brookie or mud trout, among others. A potamodromous population in Lake Superior, is known as coaster trout or, simply, as coasters. Anadromous populations which are found in coastal rivers from Long Island to Hudson Bay are sometimes referred to as salters. The brook trout is the state fish of nine U.S. states: Michigan, New Hampshire, New Jersey, New York, North Carolina, Pennsylvania, Vermont, Virginia, and West Virginia, and the Provincial Fish of Nova Scotia in Canada.
The Gila trout is a species of salmonid, related to the rainbow trout, native to the Southwest United States. Prior to 2006 the Gila trout was federally listed as endangered. In July 2006, after much work by the Game and Fish departments in New Mexico and Arizona, the US Forest Service and the US Fish and Wildlife Service, the Gila trout was down-listed to threatened, with a special provision called a "4d rule" that will allow limited sport fishing – for the first time in nearly half a century. By the time the Gila trout was closed to fishing in the 1950s, its numbers and range were so depleted and so reduced this copper-colored trout simply wasn't very accessible to anglers. As of 2011 there is fishing in both states for this fish.
The cutthroat trout(Oncorhynchus clarkii) is a fish species of the family Salmonidae native to cold-water tributaries of the Pacific Ocean, Rocky Mountains, and Great Basin in North America. As a member of the genus Oncorhynchus, it is one of the Pacific trout, a group that includes the widely distributed rainbow trout. Cutthroat trout are popular gamefish, especially among anglers who enjoy fly fishing. The common name "cutthroat" refers to the distinctive red coloration on the underside of the lower jaw. The specific name clarkii was given to honor explorer William Clark, coleader of the Lewis and Clark Expedition.
The Arctic char or Arctic charr is a cold-water fish in the family Salmonidae, native to alpine lakes, as well as Arctic and subarctic coastal waters in the Holarctic.
Sea trout is the common name usually applied to anadromous (sea-run) forms of brown trout, and is often referred to as Salmo trutta morpha trutta. Other names for anadromous brown trout are bull trout, sewin (Wales), peel or peal, mort, finnock (Scotland), white trout (Ireland), Dollaghan and salmon trout (culinary).
Salmo trutta fario, sometimes called the river trout, is a river-dwelling freshwater predatory fish from the genus Salmo of the family Salmonidae. It is one of the three main subspecies or morphs of the brown trout, besides sea trout and the lacustrine trout.
Barrow Gurney Reservoirs are three artificial reservoirs for drinking water near the village of Barrow Gurney, which lies southwest of Bristol, England. They are known by their numbers rather than names.
Chew Magna Reservoir is a 5-acre (2 ha) reservoir on the western outskirts of the village of Chew Magna, Somerset, England. It lies just north of the B3130 Winford Road.
The gillaroo is a species of trout which eats primarily snails and is only proven to inhabit Lough Melvin in Ireland.
The westslope cutthroat trout, also known as the black-spotted trout, common cutthroat trout and red-throated trout is a subspecies of the cutthroat trout and is a freshwater fish in the salmon family of order Salmoniformes. The cutthroat is the Montana state fish. This subspecies is a species of concern in its Montana and British Columbia ranges and is considered threatened in its native range in Alberta.
Dollaghan are a variety of brown trout native to Lough Neagh, Northern Ireland, and many of its tributaries. They are a potamodromous migratory trout spending much of the year in the lough, returning to the rivers in autumn to spawn. Dollaghan are much sought after by anglers in County Antrim, County Tyrone and County Londonderry due to their greater size in comparison to the non-migratory trout found in streams such as the Ballinderry River, Six Mile Water, Moyola River and River Main. They are often caught in the dark using methods very similar to that of fishing for sea trout. Many anglers regard them as an elusive species and call them 'sea trout of Lough Neagh'. Their weight varies greatly - from small fish of around 1/2 lb to large specimens of around 20 lb. There are four strains of brown trout in Ireland: Dollaghan, Gillaroo, Sonaghen and Ferox. The Dollaghan is thought to have evolved from sea trout which were land locked many years ago.
The coastal cutthroat trout, also known as the sea-run cutthroat trout, blue-back trout or harvest trout, is one of the several subspecies of cutthroat trout found in Western North America. The coastal cutthroat trout occurs in four distinct forms. A semi-anadromous or sea-run form is the most well known. Freshwater forms occur in both large and small rivers and streams and lake environments. The native range of the coastal cutthroat trout extends south from the southern coastline of the Kenai Peninsula in Alaska to the Eel River in Northern California. Coastal cutthroat trout are resident in tributary streams and rivers of the Pacific basin and are rarely found more than 100 miles (160 km) from the ocean.
The Albano is a stream of Lombardy, Italy which flows through the province of Como. It rises from the Bocchetta di Sommafiume, in the commune of Germasino and runs eastwards through the Albano valley entering Lake Como at Dongo. Between the communes of Germasino and Dongo Albano passes through Garzeno.
Angling in Yellowstone National Park is a major reason many visitors come to the park each year and since it was created in 1872, the park has drawn anglers from around the world to fish its waters. In 2006, over 50,000 park fishing permits were issued to visitors. The park contains hundreds of miles of accessible, high-quality trout rivers containing wild trout populations—over 200 creeks, streams and rivers are fishable. There are 45 fishable lakes and several large lakes are easily accessible to visitors. Additionally, the park's remote sections provide anglers ample opportunity to visit rivers, streams, creeks and lakes that receive little angling pressure. With the exception of one specially designated drainage, all the park's waters are restricted to artificial lures and fly fishing. The Madison, Firehole and a section of the Gibbon rivers are restricted to fly fishing only.
The American Angler's BookEmbracing the Natural History of Sporting Fish and the Art of Taking Them with Instructions in Fly-Fishing, Fly-Making, and Rod-Making and Directions for Fish-Breeding, to which is appended Dies Piscatoriae Describing Noted Fishing-Places, and The Pleasure of Solitary Fly-Fishing is an early American angling book by Thaddeus Norris (1811-1877) first published in 1864. Norris was known as Uncle Thad and commonly referred to in American angling history as "The American Walton".
↑ Behnke, Robert J.; Williams, Ted (2007). "Brown Trout-Winter 1986". About Trout: The Best of Robert J. Behnke from Trout Magazine. Guilford, CT: Globe Pequot. p.45. ISBN978-1-59921-203-6.
1 2 3 4 Behnke, Robert J.; Williams, Ted (2007). "Brown Trout-Winter 1986". About Trout: The Best of Robert J. Behnke from Trout Magazine. Guilford, CT: Globe Pequot. pp.45–50. ISBN978-1-59921-203-6.
↑ Newton, Chris (2013). "The Trout in India". The Trout's Tale – The Fish That Conquered an Empire. Ellesmere, Shropshire: Medlar Press. pp.79–95. ISBN978-1-907110-44-3.
↑ Newton, Chris (2013). "The Monsters of Kerguelen". The Trout's Tale – The Fish That Conquered an Empire. Ellesmere, Shropshire: Medlar Press. pp.161–170. ISBN978-1-907110-44-3.
↑ Sánchez-Hernández, J., Servia, M.J., Vieira-Lanero, R. & Cobo F. (2013). Ontogenetic dietary shifts in a predatory freshwater fish species: the brown trout as an example of a dynamic fish species. In: New Advances and Contributions to Fish Biology, Hakan Türker (Ed.). ISBN978-953-51-0909-9, InTech, Croatia, 271–298 pp.
↑ Jensen, H.; Kiljunen, M.; Amundsen, P-A. (2012). "Dietary ontogeny and niche shift to piscivory in lacustrine brown trout Salmo trutta revealed by stomach content and stable isotope analyses". Journal of Fish Biology. 80 (7): 2448–2462. Bibcode:2012JFBio..80.2448J. doi:10.1111/j.1095-8649.2012.03294.x. PMID22650427.
↑ Sánchez-Hernández, J.; Vieira-Lanero, R.; Servia, M.J.; Cobo, F. (2011a). "First feeding diet of young brown trout fry in a temperate area: disentangling constrains and food selection". Hydrobiologia. 663 (1): 109–119. doi:10.1007/s10750-010-0582-3. S2CID23870995.
Marco-Rius, Francisco; Sotelo, Graciela; Caballero, Pablo; Morán, Paloma (2013). "Insights for planning an effective stocking program in anadromous brown trout (Salmo trutta)". Canadian Journal of Fisheries and Aquatic Sciences. 70 (7): 1092–1100. doi:10.1139/cjfas-2013-0084..
↑ Herd, Andrew Dr (2001). "Beginnings". The Fly. Ellesmere, Shropshire: Medlar Press. pp.19–74. ISBN1-899600-19-1.
↑ Newton, Chris (2013). "Two Fish in One". The Trout's Tale – The Fish That Conquered an Empire. Ellesmere, Shropshire: Medlar Press. pp.31–36. ISBN978-1-907110-44-3.
↑ Newton, Chris (2013). "Falklands' Silver". The Trout's Tale – The Fish That Conquered an Empire. Ellesmere, Shropshire: Medlar Press. pp.149–159. ISBN978-1-907110-44-3. The Chartres produces some great fishing ...When he was reunited with the party a few hours later, he had taken 15 sea trout from 7 lbs to 14 1/2 lbs – world-class fishing by any yardstick
J.L. Bagliniere; G. Maisse; J. Watson (1999). Biology and Ecology of the Brown Sea Trout. Springer Praxis Books. ISBN1852331178.
Elliot, J.M. (1994). Quantitative Ecology and the Brown Trout. Oxford, UK: Oxford University Press. ISBN0198540906.
Newton, Chris (2013). The Trout's Tale – The Fish That Conquered an Empire. Ellesmere, Shropshire: Medlar Press. ISBN978-1-907110-44-3.
Marston, R.B. (Summer 1985). "Brown Trout (Salmo fario)"(PDF). The American Fly Fisher. Manchester, VT: American Museum of Fly Fishing. 12 (3): 7–8. Archived from the original(PDF) on 2015-07-01. Retrieved 2014-11-19.
External links
Wikimedia Commons has media related to Salmo trutta.
This page is based on this Wikipedia article Text is available under the CC BY-SA 4.0 license; additional terms may apply. Images, videos and audio are available under their respective licenses.
