Galaxiidae Temporal range: Maastrichtian occurrence Possible | |
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A mountain galaxias ( Galaxias olidus ) | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Superorder: | Osmeromorpha |
Order: | Galaxiiformes |
Family: | Galaxiidae Bonaparte 1832 |
Genera | |
See text |
The Galaxiidae are a family of mostly small freshwater fish in the Southern Hemisphere. The majority live in Southern Australia or New Zealand, but some are found in South Africa, southern South America, Lord Howe Island, New Caledonia, and the Falkland Islands. One galaxiid species, the common galaxias (Galaxias maculatus), is probably the most widely naturally distributed freshwater fish in the Southern Hemisphere. They are coolwater species, found in temperate latitudes, with only one species known from subtropical habitats. [1] Many specialise in living in cold, high-altitude upland rivers, streams, and lakes.
Some galaxiids live in fresh water all their lives, but many have a partially marine lifecycle. In these cases, larvae are hatched in a river, but are washed downstream to the ocean, later returning to rivers as juveniles to complete their development to full adulthood. This pattern differs from that of salmon, which only return to fresh water to breed, and is described as amphidromous. [2]
Freshwater galaxiid species are gravely threatened by exotic salmonid species, particularly trout species, which prey upon galaxiids and compete with them for food. Exotic salmonids have been recklessly introduced to many different land masses (e.g. Australia, New Zealand), with no thought as to impacts on native fish, or attempts to preserve salmonid-free habitats for them. Numerous localised extinctions of galaxiid species have been caused by the introduction of exotic salmonids, and a number of freshwater galaxiid species are threatened with overall extinction by exotic salmonids. [1]
Phylogenetic evidence alternatively places galaxiids within the Protacanthopterygii, or more recently as the sister group to the Neoteleostei. Their ancestors are thought to have diverged from the neoteleosts around the Triassic-Jurassic boundary. [3] [4]
The earliest definitive fossils of galaxiids are from the Miocene of New Zealand, which can be placed in the extant genus Galaxias . This young fossil range contrasts with the presumed ancient origins of the group. In 1998, a possible Late Cretaceous (Maastrichtian) galaxiid from South Africa was described as Stompooria . [5] However, later studies have questioned this assignment, as Stompooria differs from galaxiids in many morphological traits, especially in the presence of scales, although it being an ancestral galaxiid that had not yet developed galaxiid traits could not be ruled out. [6] Other taxonomic treatments have instead placed Stompooria as part of an extinct clade sister to the Esociformes and Salmoniformes. [4]
About 50 species are in the family Galaxiidae, grouped into seven genera: [7]
Galaxiids are found around the south eastern seaboard of Australia and in some parts of south western Australia. The galaxiids and the temperate perches (Percichthyidae) are the dominant native freshwater fish families of southern Australia. Species common to all areas include:
South east Australian mainland
Threatened species are:
Western Australia
Tasmania Seventeen species of galaxiids have been found in Tasmania. The most common species are:
Tasmanian endangered species include:
Twenty-three species of galaxiids have been discovered in New Zealand, and prior to the introduction of non-native species such as trout, they were the dominant freshwater fish family. Most of these live in fresh water all their lives. However, the larvae of five species of the genus Galaxias develop in the ocean, where they form part of the zooplankton and return to rivers and streams as juveniles (whitebait), where they develop and remain as adults. All Galaxias species found in New Zealand are endemic, except for Galaxias brevipinnis (koaro) and Galaxias maculatus (inanga).
The juveniles of those galaxiids that develop in the ocean and then move into rivers for their adult lives are caught as whitebait while moving upstream and are much valued as a delicacy. Adult galaxiids may be caught for food, but they are generally not large. In some cases, their exploitation may be banned (e.g. New Zealand) unless available to indigenous tribes.
In addition to serious impacts from exotic trout species, Australian adult galaxiids suffer a disregard from anglers for being "too small" and "not being trout". This is despite the fact that several Australian galaxiid species, though smallish, grow to a sufficient size to be catchable and readily take wet and dry flies, and that one of these species — the spotted galaxias — was keenly fished for in Australia before the introduction of exotic trout species. A handful of fly-fishing exponents in Australia are rediscovering the pleasure of catching (and releasing) these Australian native fish on ultralight fly-fishing tackle.
The Australian grayling is a primarily freshwater fish found in coastal rivers in south-eastern mainland Australia and Tasmania. In past decades it has also been known as the cucumber mullet or cucumber herring, for its cucumber-like odour.
The climbing galaxias or kōaro is a fish of the family Galaxiidae found in Australia, New Zealand, and nearby islands. The name climbing galaxias is used in Australia, and koaro or kōaro in New Zealand. Further vernacular names include short-finned galaxias, broad-finned galaxias, Cox's mountain galaxias, and Pieman galaxias.
The common galaxias or inanga is a very widespread Southern Hemisphere fish in the family Galaxiidae. It is a slim, narrow fish with a forked tail and a mottled, spotty pattern, typically about 10 cm (4 in) long when fully grown. It lives in fresh water, but spawns at river mouths and spends the first six months of its life at sea, returning en masse in spring. Its vernacular names include cowfish, jollytail, common jollytail, eel gudgeon, inaka, native trout, pulangi, puye, slippery tarki, spotted minnow, Falklands minnow and whitebait.
The Chatham mudfish, formerly known as the Chathams galaxias, is a galaxiid fish endemic to two small, peaty lakes in southern Chatham Island, New Zealand.
Neochanna is a genus of galaxiid fishes, commonly known as mudfish, which are native to New Zealand and south-eastern Australia.
The Tasmanian mudfish, Neochanna cleaveri, is a small Australian amphidromous fish in the galaxiid family, of the order Osmeriformes.
The banded kōkopu is a galaxiid of the genus Galaxias, found only in New Zealand, including the Chatham and Stewart/Rakiura islands. It commonly grows to 20–25 cm, but has been recorded growing to around 30 cm. Juvenile banded kōkopu are good climbers and can climb up waterfalls and other vertical surfaces by moving into the splash zone and wriggling up the surface, using the water surface tension and their large downturned fins for grip.
The shortjaw kōkopu is a galaxiid fish, endemic to New Zealand. They are large, scaleless fish which inhabit stable pools in cascading, bouldery streams with forest cover. Shortjaw kōkopu are amphidromous, with the fry going to sea after hatching, and returning as juveniles to fresh water where they grow to adulthood. They may grow to a maximum of 350 mm (14 in) in total length, but more commonly reach 150–200 mm (5.9–7.9 in).
The giant kōkopu is a threatened species of ray-finned fish in the genus Galaxias, found only in New Zealand. It can reach up to 58 cm (23 in) in length and 2.7 kg (6.0 lb) in weight, making it the largest species in the family Galaxiidae. It is a mainly lowland species, commonly found in slow-flowing streams, wetlands, lakes, and lagoons. Most populations have an amphidromous life cycle, with larvae going to sea soon after hatching and returning about four months later as small juveniles, 4.5–5 cm (1.8–2.0 in). Juvenile giant kōkopu form a small part of the annual whitebait catch.
The dwarf inanga is a galaxiid of the genus Galaxias, found in the North Island of New Zealand.
Eldon's galaxias is an endangered galaxiid fish endemic to New Zealand. One of several fishes in the genus Galaxias found in Central Otago, it has a very small home range and is at risk of being driven to extinction by trout introduced for recreational fishing.
The common river galaxias or Canterbury galaxias is a galaxiid fish of the genus Galaxias, found only in Canterbury, New Zealand.
Gollum galaxias is a galaxiid of the genus Galaxias, found only in southern temperate regions in New Zealand. The species is widespread in rivers throughout the Southland Plains as well as the Von River and Nevis River tributaries of the Clutha River The species is found on Stewart Island/Rakiura, and throughout the Catlins.
Galaxias is a genus of small freshwater fish in the family Galaxiidae, and are frequently referred to as the galaxiids. These highly adaptable fish are typically found at temperate latitudes across the Southern Hemisphere.
Galaxiella is a genus of Australian fish of the family Galaxiidae.
Spotted galaxias is a largish, primarily-freshwater galaxias species found in southern Australia. Spotted galaxias are perhaps the most beautiful of the Australian galaxias species. They are a somewhat tubular, deep-bodied fish, with a dusky brownish-red colouration overlain with dark, haloed spots, dramatic black edges to dorsal, anal and pelvic fins, and a dark diagonal stripe through the eye.
Gerald Stokell was a New Zealand amateur ichthyologist.
Whitebait is a collective term for the immature fry of fish, typically between 25 and 50 millimetres long. Such young fish often travel together in schools along coasts, and move into estuaries and sometimes up rivers where they can be easily caught using fine-meshed fishing nets. Whitebaiting is the activity of catching whitebait.
Robert Montgomery McDowall was one of New Zealand's most prominent freshwater ichthyologists.
Galaxiella toourtkoourt, known as the little galaxias, is a species of fish in the family Galaxiidae. It is among the smallest galaxiid species known.