Cladistia Temporal range: Permian and Carboniferous records) (Possible | |
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Polypterus senegalus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Subclass: | Cladistia Pander 1860 emend. Cope 1871 sensu Lund 2000 |
Orders | |
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Cladistia is a subclass of bony fishes whose only living members are the bichirs of tropical Africa. [1] Their major synapomorphies are a heterocercal tail in which the dorsal fin has independent rays, and a posteriorly elongated parasphenoid.
Cladistia are the earliest diverging branch of living Actinopterygii, and are thought to have diverged from the Actinopteri, the group which includes all other living ray finned fish, by the Carboniferous. [2] However, the fossil range for the only extant order (Polypteriformes) is comparatively young, only reaching as far back as the mid-Cretaceous of South America and Africa, and the two extant genera of bichir only diverged around the Miocene. [3]
Aside from bichirs, other extinct fish groups thought to be members of the group include the Scanilepiformes, known from Triassic (and possibly Permian [4] ) of the Northern Hemisphere. [5] [2] [6] The Guildayichthyiformes of Carboniferous North America are also sometimes considered cladistians, but this is thought to be dubious, with other authorities placing them as a stem-group to Neopterygii. [6] [7]
Based on work done by Near et al (2017) and Lund (2000): [8]
Actinopterygii, members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of skin supported by radially extended thin bony spines called lepidotrichia, as opposed to the bulkier, fleshy lobed fins of the sister class Sarcopterygii. Resembling folding fans, the actinopterygian fins can easily change shape and wetted area, providing superior thrust-to-weight ratios per movement compared to sarcopterygian and chondrichthyian fins. The fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the articulation between these fins and the internal skeleton.
Bichirs and the reedfish comprise Polypteridae, a family of archaic ray-finned fishes and the only family in the order Polypteriformes.
The reedfish, ropefish, or snakefish, Erpetoichthys calabaricus, is a species of fish in the family Polypteridae alongside the bichirs. It is the only member of the genus Erpetoichthys. It is native to fresh and brackish waters in West and Central Africa. The reedfish possesses a pair of lungs in addition to gills, allowing it to survive in very oxygen-poor water. It is threatened by habitat loss through palm oil plantations, other agriculture, deforestation, and urban development.
Polypterus is a genus of freshwater fish in the bichir family (Polypteridae) of order Polypteriformes. The type species is the Nile bichir. Fish in this genus live in various areas in Africa. Polypterus is the only known vertebrate to have lungs, but no trachea.
The Palaeonisciformes, commonly known as "palaeoniscoids" are an extinct grouping of primitive ray-finned fish (Actinopterygii), spanning from the Silurian/Devonian to the Cretaceous. They are generally considered paraphyletic, but their exact relationships to living ray-finned fish are uncertain. While some and perhaps most palaeoniscoids likely belong to the stem-group of Actinopteryii, it has been suggested that some may belong to the crown group, with some of these possibly related to Cladistia and/or Chondrostei. Many palaeoniscoids share a conservative body shape and a similar arrangement of skull bones, though paleoniscoids as a whole exhibit considerable diversity in body shape.
Birgeria is a genus of carnivorous marine ray-finned fish from the Triassic period. Birgeria had a global distribution, with fossil known from Madagascar, Spitsbergen, Germany, Switzerland, Italy, Slovenia, China, Russia, Canada and Nevada, United States. The oldest fossils are from Griesbachian aged beds of the Wordie Creek Formation of East Greenland. Birgeria existed throughout the entire Triassic period, from the very beginning just after the Permian-Triassic mass extinction, up to the very end with its extinction during the Triassic-Jurassic mass extinction.
Polypterus bichir, the Nile bichir, is a fish which lives in the Nile and some of its tributaries in Africa. It is a dark grayish color on the top, with a dark vertical marking and bands on the flank. This marking is more prominent on juveniles, and fades as the fish grows.
The barred bichir, armoured bichir, bandback bichir, or banded bichir is a species of fish belonging to the class Actinopterygii, or the ray-finned fish, which constitute an order of the class of the bony fish. an elongated fish found in the Congo River, specifically in the upper and middle portions. This species is one of the more commonly available in commercial pet stores.
Scanilepis is an extinct genus of prehistoric bony fish that lived during the Rhaetian-?Hettangian ages. The type species, S. dubia, is known from the Rhaetian freshwater deposits of the Bjuv member of the Höganäs Formation, southwestern Sweden. A second species, S. spitzbergensis was mentioned from the Hettangian of the Festning section of the Grøfjorden area in Spitsbergen, Norway but was never described.
Boreolepis is an extinct genus of prehistoric marine ray-finned fish that lived from the Late Kungurian to the Wuchiapingian of the Permian period. It inhabited the high northern latitudes in what is now Greenland and European Russia.
Saurichthyiformes is an extinct order of ray-finned fish which existed in Asia, Africa, Australia, Europe and North America, during the late Permian to early Middle Jurassic. Saurichthyiiformes comprise two families, Saurichthyidae and Yelangichthyidae. Yelangichthyidae is monotypic, containing only the genus Yelangichthys. The gar or needlefish-like Saurichthyidae is primarily known from the genus Saurichthys. Additionally, the subgenera SaurorhynchusCostasaurichthys, Eosaurichthys, Lepidosaurichthys, and Sinosaurichthys are frequently used to group species, and are sometimes considered separate genera. Species are known from both marine end freshwater deposits. They had their highest diversity during the Early and Middle Triassic. Their phylogenetic position is uncertain, while they have often been considered members of Chondrostei, and thus related to living sturgeons and paddlefish, phylogenetic analysis of well-preserved remains has considered this relationship equivocal. They may actually belong to the stem-group of Actinopterygii, and thus not closely related to any living group of ray-finned fish.
Beishanichthys is an extinct genus of freshwater ray-finned fish which existed in Gansu Province, China during the Olenekian age of the early Triassic period. It contains a single species, B. brevicaudalis, first named by Guang-Hui Xu, Ke-Qin Gao in 2011 based on fossils from the Lower Triassic lake deposits exposed in northern Gansu, China. It is considered a scanilepiform, a group of early cladistians related to the modern bichirs, although Beishanichthys was not incorporated into the analyses that found this phylogenetic placement among the cladistians.
Bawitius is an extinct genus of giant polypterid from the Upper Cretaceous Bahariya Formation of Egypt. The type species is B. bartheli, named as a species of Polypterus in 1984, and the genus etymology comes from Bawiti, the principal settlement of the Bahariya Oasis in Egypt. It is known from the holotype TU-B SFB 69 Vb 003 : left ectopterygoid scales and some sparse scales.
Polypterus palmas, also called the shortfin or marbled bichir, is a fish in the family Polypteridae found in freshwater environments throughout West Africa.
Guildayichthyidae is a prehistoric family of marine fish from the Mississippian Bear Gulch Limestone of Montana. It is the only family in the order Guildayichthyiformes. Guildayichthyids possess an uncommon mixture of primitive and modern characteristics in their skull bones.
Discoserra is a prehistoric ray-finned fish from the Mississippian of the Bear Gulch Limestone in Montana. It is a member of the Guildayichthyiformes with a round body and a skull possessing primitive and modern traits. Discoserra is about 60 mm long. In 2006, Discoserra was hypothesized to be a stem neopterygian, although it has alternatively been placed in Cladistia along with other Guildayichthyiformes.
Polypterus mokelembembe is a species of the fish genus Polypterus, found in the central basin of the Congo River. It was once considered a morph of the closely related Polypterus retropinnis, but was given species status in 2006 with a description that reclassified both fishes. Because of the recency of the species' description and the fact that P. mokelembembe is the paralectotype of P. retropinnis, they are often mistaken for one another in the aquarium trade.
Serenoichthys is an extinct genus of small bichir from the Late Cretaceous (Cenomanian) of southeastern Morocco. The genus is monotypic, the type and only species being Serenoichthys kemkemensis. Only known at first from postcranial skeletons, complete specimens were later discovered.
Acropholis is an extinct genus of prehistoric marine bony fish that lived during the Wuchiapingian age in what is now Greenland and Hesse (Germany).
Tanaocrossus is an extinct genus of primitive freshwater ray-finned fish that inhabited southwestern and eastern North America during the Late Triassic period. It contains a single known species, T. kalliokoskii, known from the United States, with indeterminate species also known.