List of acanthodian genera

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Various primitive acanthodians from Early Devonian England and Scotland, Mesacanthus pusillus, Parexus falcatus, Ishnacanthus gracilis. Mesacanthus Parexus Ischnacanthus.JPG
Various primitive acanthodians from Early Devonian England and Scotland, Mesacanthus pusillus , Parexus falcatus , Ishnacanthus gracilis.

This list of acanthodian genera is an attempt to create a comprehensive listing of all genera that have ever been included in the subclass Acanthodii, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful ( nomina dubia ), or were not formally published ( nomina nuda ), as well as junior synonyms of more established names, and genera that are no longer considered acanthodians. Not counting their descendants, the modern chondricthyans, all acanthodians are extinct. Extinct genera are marked with a dagger ()

Contents

Naming conventions and terminology

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

The List

GenusStatusAgeLocationNotesImages
Acanthacanthus [1] Valid.Early Lochkovian (earliest Devonian). [1] Severnaya Zemlya, Russia. [1] A vesperaliid based on scales. [2]
Acanthodopsis Valid. Carboniferous. [3] Europe, Australia. [3] An unusual acanthodid with tooth-like ossifications on its jaws. [3]
Acanthopora [1] Valid.Early Lochkovian (earliest Devonian). [1] Severnaya Zemlya, Russia. [1] A small ischnacanthid similar to Poracanthodes . [1]
Acanthospina [1] Valid.Early Lochkovian (earliest Devonian). [1] Severnaya Zemlya, Russia. [1] A probable ischnacanthiform with scales similar to hybodontiform sharks. [1] [3]
Acanthodes Valid. Devonian? to Cisuralian (Early Permian). [4] Europe, North America, China?One of the most well-known acanthodians, a filter-feeding acanthodid acanthodiform. Acanthodes lopatini.png
Acritolepis [1] Valid.Early Lochkovian (earliest Devonian). [1] Severnaya Zemlya, Russia. [1] Namesake of the ischnacanthiform family Acritolepidae. [2] [5]
Aganacanthus Valid or junior synonym. Mississippian / Early Carboniferous. Scotland.A possible gyracanthid known from a single worn fin spine, may be an indeterminate gnathostome or a junior synonym of Gyracanthus . [6]
Ankylacanthus [7] Valid. Emsian (Early Devonian). [7] Canada. [7] An acanthodian based on fin spines, possibly one of the oldest gyracanthids.
Antacanthus Nomen dubium? Mississippian / Early Carboniferous. [8] Belgium. [8] A possible gyracanthid known from a single fin spine. [9] [6]
Antarctonchus [10] Valid. Givetian (Middle Devonian). [10] Antarctica. [10] An Antarctic acanthodian based on fin spine casts. [10]
Apateacanthus Valid. Late Devonian. [3] New York, U.S.A. [3] Based on a fin spine with large denticles, sometimes mistaken for an ischnacanthiform jaw. [3]
Archaeacanthus Valid. Eifelian to Givetian (Middle Devonian). [11] Europe. [11] Based on fin spines common in Middle Devonian strata of Siberia and Northeastern Europe. [11]
Arcticacanthus [1] Valid.Early Lochkovian to Emsian (Early Devonian). [1] Severnaya Zemlya, Russia. [1] A possible ischnacanthid based on scales. [1]
Arenaceacanthus [12] Valid. Wenlock to early Pridoli (Silurian). [13] [12] Lithuania. [12] A probable ischnacanthid based on scales, common in Lithuania. [12]
Atopacanthus Valid. Eifelian to Early Famennian (Middle Devonian to Late Devonian). [14] [15] North America, Europe, [14] Iran. [15] A widespread and long-lived ischnacanthid. [14]
Brachyacanthus Valid. Lochkovian (Early Devonian). [16] Scotland. [17] A small climatiid [16] from the Lower Old Red Sandstone of Scotland. [17]
Bracteatacanthus [12] Valid.Latest Pridoli (latest Silurian). [13] [12] Lithuania. [12] A probable ischnacanthid based on scales. [12]
Brochoadmones Valid. Lochkovian (Early Devonian). [18] Canada. [18] A deep-bodied and well-preserved "climatiiform" from the Man-On-The-Hill (MOTH) locality in the MacKenzie Mountains of Canada. [18] BrochoadmonesDB15.jpg
Bryantonchus [19] Valid. Emsian (Early Devonian). [19] U.S.A. [19] A probable diplacanthiform based on fin spines. [19]
Byssacanthoides Valid. Givetian (Middle Devonian). [10] Antarctica. [10] An Antarctic acanthodian based on fin spine casts. [10]
Cacheacanthus [19] Valid. Emsian (Early Devonian). [19] U.S.A. [19] Based on ischnacanthid jaw fragments from the western United States. [19]
Campylodus Nomen dubium? Ludlow [20] to Lochkovian? [21] (late Silurian to earliest Devonian)? Europe. [20] [21] Jaw fragments probably belonging to indeterminate ischnacanthiforms. [20] [21]
Canadalepis Valid. Lochkovian? [22] to Emsian (Early Devonian). [23] [24] Asia, [23] Australia, [24] Canada. [22] A possible climatiid based on scales. [23] [24] [22]
Carycinacanthus Junior synonym. [4] Tournaisian (earliest Mississippian / Carboniferous). [4] Russia. [4] A junior synonym of Acanthodes lopatini . [4]
Cassidiceps [25] Valid. Lochkovian (Early Devonian). [25] Canada. [25] A toothless, deep-bodied MOTH locality acanthodian with complex head armor. [25] Probably closely related to the acanthodiforms. [26]
Cheiracanthoides Valid. Pridoli to Frasnian (latest Silurian to Late Devonian). [13] [27] Worldwide.A widespread and long-lasting possible climatiid based on scales. [27]
Cheiracanthus Valid. Middle Devonian. [28] Europe. [28] The fairly common namesake of the acanthodiform family Cheiracanthidae. [28] Cheiracanthus murchisoni life restoration.jpg
Climatius Valid. Lochkovian (Early Devonian). [29] U.K., [29] [30] Portugal. [21] The namesake of the family Climatiidae. [29] Climatius BW.jpg
Culmacanthus Valid. Givetian to Frasnian (Middle Devonian to Late Devonian). [31] [32] Australia, [31] [32] Antarctica. [10] A distinctive diplacanthiform with reduced intermediate spines and large cheek plates. [31] [32]
Devononchus Valid. Givetian [11] to Famennian [33] (Middle Devonian to Late Devonian). Europe.Based on fin spines and scales common in the areas corresponding to Baltica.
Diplacanthus Valid. Middle Devonian [34] to Late Devonian. [35] Scotland, [34] Russia, [1] South Africa, [35] Canada? [34] The widespread, long-lasting, and fairly common namesake of the family Diplacanthidae and order Diplacanthiformes. [34] Diplacanthus.png
Doliodus Valid (non-acanthodian). Emsian (Early Devonian). [36] Canada. [37] [36] A spiny stem-chondrichthyan which acts as a transitional form between acanthodians and non-acanthodian traditional chondrichthyans. [37] [38] Doliodus22DB.jpg
Eifellepis Valid. Emsian to Eifelian (Early Devonian to Middle Devonian). [22] Germany, Belgium. [22] An indeterminate acanthodian based on scales. [2]
Erriwacanthus Valid. Pragian (Early Devonian). [39] U.K., Ukraine. [40] A climatiid with large and complex spines on its shoulder girdle. [40]
Endemolepis Valid. Pridoli to Lochkovian (latest Silurian to Early Devonian). [12] Europe. [12] A possible climatiid based on scales. [12]
Erymnacanthus [41] Valid. Lochkovian (Early Devonian). [41] Canada. [41] A medium-sized MOTH locality ischnacanthid. [41] Erymnacanthus clivus jaw cast.jpg

Erymnacanthus clivus jaw cast

Eupleurogmus Nomen dubium. [42] Mississippian / Early Carboniferous? [42] Australia. [42] A dubious taxon based on Acanthodes -like scales. [42]
Euryacanthus [41] Valid. Lochkovian (Early Devonian). [41] Canada. [41] A medium-sized MOTH locality ischnacanthid. [41]
Euthacanthus Valid.Early Lochkovian (earliest Devonian). [30] U.K. [30] A slender acanthodian of uncertain affinities, despite its well-described anatomy. [43] Euthacanthus spp.jpg
Fallodentus [44] Valid. Givetian (Middle Devonian). [44] Scotland. [44] A cheiracanthid acanthodiform closely related to Homalacanthus . [44]
Fanjingshania [45] Valid. Aeronian (Early Silurian, Llandovery Epoch). [45] China. [45] A climatiiform, one of the earliest known acanthodean genus. [45] Fanjingshania.jpg

Fanjingshania fin spine

Fecundosquama [12] Valid. Pridoli (late Silurian). [13] Lithuania. [12] A tchunacanthid [2] based on scales. [12]
Florestacanthus [46] Valid.Late Frasnian to early Famennian (Late Devonian). [46] Colombia. [46] A diplacanthid diplacanthiform and one of the few acanthodians endemic to northwestern Gondwana (now South America). [46]
Gemuendolepis Valid? Early Devonian. [47] Germany. [48] A poorly-described taxon based on scales.
Ginkgolepis [49] Valid. Eifelian to Famennian? (Middle Devonian to Late Devonian). [49] Estonia, Russia. [49] A cheiracanthid acanthodiform based on scales. [49]
Gladbachus Valid (non-acanthodian?). Givetian (Middle Devonian). [50] Germany. [50] A large filter-feeding stem-chondrichthyan, sometimes positioned among acanthodians despite its very dissimilar anatomy. [50] Gladbachus adentatus.jpg
Gladiobranchus Junior synonym? [51] Lochkovian (Early Devonian). [52] Canada. [52] A well-preserved diplacanthiform which may be synonymous with Uraniacanthus curtus . [51]
Gomphacanthus [53] Nomen dubium? Carboniferous. England. [53] A possible gyracanthid based on a worn fin spine. [54]
Gomphonchoporus Valid. Pridoli to Lochkovian (late Silurian to Early Devonian). [55] Europe, [55] Australia, [56] Canada, [57] Greenland. [39] A widespread poracanthodid based on scales. [55]
Gomphonchus Valid. Ludlow to Emsian? (late Silurian to Early Devonian). [55] [58] Europe, [55] Australia?, China? [58] A widespread ischnacanthid primarily based on scales. [55]
Granulacanthus [59] Valid.Late Wenlock or early Ludlow (Silurian). [59] Canada. [59] A possible ischnacanthiform [20] based on distinctive fin spines. [59]
Grenfellacanthus [60] Valid.Latest Famennian (latest Devonian). [60] Australia. [60] One of the youngest and largest ischnacanthids, nearly a meter in length. [60]
Gyracanthides Valid. Pragian to Visean (Early Devonian to Mississippian). [6] Worldwide. [6] A long-lasting and widespread gyracanthid with flattened fin spines. [9] [6] Gyracanthides murrayi.png
Gyracanthus Valid. Lochkovian to Moscovian (Early Devonian to Pennsylvanian) [6] Europe, North America, Bolivia? [6] The long-lasting namesake of the family Gyracanthidae. [6] Gyracanthus NT small cropped.png
Hanilepis Valid.Late Ludlow (late Silurian). [61] China. [61] An indeterminate acanthodian based on scales. [61]
Halimacanthodes [62] Valid. Frasnian (Late Devonian). [62] Australia. [62] A howittacanthid acanthodiform, the only acanthodian known from the Gogo Formation lagerstätte. [62]
Haplacanthus Valid. Eifelian to early Famennian (Middle Devonian to Late Devonian). [11] [33] Europe, Greenland. [11] A cheiracanthid [44] acanthodiform based on fin spines common in the areas corresponding to Baltica. [11]
Helenacanthus Valid? (non-acanthodian) [3] Emsian (Early Devonian). [19] Wyoming, U.S.A. [3] Has historically been considered acanthodian material, but probably armor from an actinolepid placoderm instead. [3]
Helolepis Junior synonym. Emsian to Eifelian (Early Devonian to Middle Devonian). [24] Ohio, U.S.A. [24] A junior synonym of Cheiracanthoides comptus . [63]
Homacanthus Junior synonym? Eifelian (Middle Devonian) [11] to Pennsylvanian? [37] Europe, [11] North America? [37] An acanthodian based on fin spines, though many of its species and specimens have been reassigned to other taxa. [34] [37]
Homalacanthus Valid. Frasnian (Late Devonian). [64] Canada. [64] A cheiracanthid [44] acanthodiform known to have played a large role in the food chain of the Miguasha Lagerstätte. [64]
Howittacanthus [42] Valid. Frasnian (Late Devonian). [42] Australia. [42] The abundant namesake of the acanthodiform family Howittacanthidae. [65] [62]
Iranolepis [66] Valid.Early Frasnian (Late Devonian). [66] Iran. [66] A possible climatiid based on scales. [66]
Ischnacanthus Valid. Lochkovian (Early Devonian). [67] U.K., [30] [67] Portugal. [21] The abundant namesake of the ischnacanthiforms. [67] Ischnacanthus gracilis.png
Kathemacanthus [25] Valid. Lochkovian (Early Devonian). [25] Canada. [25] A Brochoadmones -like acanthodian from the MOTH locality, with a "necklace" of artichoke-shaped scales similar to those of chondrichthyans. [25] [68]
Laliacanthus Valid. Emsian to Eifelian (Early Devonian to Middle Devonian). [22] [11] Europe, [11] Saudi Arabia. [22] A possible climatiid based on scales. [22]
Latviacanthus [69] Valid."upper Lower Devonian" [69] Latvia. [69] A possible climatiid with large ischnacanthiform-like jaw plates. [69] [18]
Lenacanthus Valid.Early Llandovery (earliest Silurian). [2] Russia. [2] A tchunacanthid based on scales, one of the earliest acanthodians. [2]
Lijiangichthys Valid. Pragian (Early Devonian). [23] China. [23] An indeterminate acanthodian based on scales. [23]
Lodeacanthus Valid. Frasnian (Late Devonian). [70] Latvia. [70] A mesacanthid acanthodiform known from a complete growth series. [70] [71]
Lupopsyrus Valid. Lochkovian (Early Devonian). [26] Canada. [26] A MOTH locality acanthodian with two rows of enlarged scales on either side of its body. [26]
Machaeracanthus Valid.Late Silurian to Eifelian (Middle Devonian). [72] Worldwide. [72] A widespread possible ischnacanthiform which occupies its own monotypic family. [72] An examination of the Devonian fishes of Michigan fig-11-full.png

Machaeracanthus fin spines

Machaeraporus [57] Valid. Pridoli to Lochkovian (late Silurian to Early Devonian). [57] Canada, Europe. [73] [57] A poracanthodid ischnacanthiform based on scales. [57]
Markacanthus Valid. Eifelian to Givetian (Middle Devonian). [44] Belarus, Scotland. [44] A fairly typical cheiracanthid acanthodiform. [44]
Marsdenius Valid.Mississippian / Early Carboniferous. [3] England. [3] An obscure possible ischnacanthid based on a specimen which is now lost. [3] [59]
Melanoacanthus [71] Valid. Emsian (Early Devonian). [71] Canada. [71] A small mesacanthid acanthodiform with broad head plates. [71]
Mesacanthus Valid. Early Devonian to Middle Devonian. [74] Scotland. [74] The small but fairly common namesake of the acanthodiform family Mesacanthidae. [74] Mesacanthus.jpg
Milesacanthus [10] Valid. Pragian to Frasnian (Early Devonian to Late Devonian). [10] [66] Worldwide. [22] A widespread diplacanthid diplacanthiform. [10] [22]
Minioracanthus Valid. Eifelian to Givetian (Middle Devonian). [11] Europe. [11] An indeterminate acanthodian based on scales. [2]
Monopleurodus Valid. Pridoli (late Silurian). Estonia. [75] Poorly-known.
Monospina Valid.Late Pridoli (latest Silurian). [13] Latvia, Lithuania. [13] An acritolepid ischnacanthiform based on scales. [2]
Nerepisacanthus [20] Valid. Ludlow to late Pridoli (late Silurian). [20] [5] Canada. [5] An acritolepid ischnacanthiform and one of the most well-preserved Silurian acanthodians. [5] Nerepisacanthus.png
Nobilesquama [2] Valid. Lochkovian to Emsian (Early Devonian). [2] Russia. [2] An indeterminate acanthodian based on scales. [2]
Nodocosta Valid. Emsian to Frasnian (Early Devonian to Late Devonian). [19] Europe, U.S.A. [19] A possible climatiid based on fin spines. [19]
Nodonchus Junior synonym? Lochkovian (Early Devonian). England, [30] Saudi Arabia? [22] Based on fin spines, but may be a junior synonym of Parexus . [30]
Nostolepis Valid. Wenlock to early Frasnian (mid-Silurian to Late Devonian). [13] [33] Worldwide. [2] One of the most widespread, long-lasting, and abundant acanthodians based primarily on scales. [2]
Nostovicina [2] Valid. Pridoli to Emsian (late Silurian to Early Devonian). [2] Worldwide. [2] A widespread indeterminate acanthodian based on scales and fin spines. [2]
Obruchevacanthus [76] Valid.Late Lochkovian (Early Devonian). [76] Spain. [76] A poracanthodid ischnacanthiform closely related to Trundlelepis . [76]
Obtusacanthus Valid. Lochkovian (Early Devonian). [77] Canada. [77] An unusual MOTH locality gnathostome, likely an acanthodian closely related to true chondrichthyans. Despite lacking teeth, it was a predator with specialized lip scales. [77]
Onchus Valid.Late Llandovery? [78] to Famennian (early Silurian to Late Devonian). [79] Europe, North America.A possible ischnacanthiform based on fin spines; may be one of the oldest known acanthodians. [3]
Oracanthus Valid? Mississippian. Europe, North America.A poorly-known gyracanthid based on fin spines. [9] [6]
Orcadacanthus [80] Valid. Middle Devonian. Scotland.A small mesacanthid based on species formerly placed in Acanthodes and Mesacanthus . [80]
Paranostolepis Valid. Late Devonian. Canada.A poorly-known indeterminate acanthodian based on scales. [2]
Parexus Valid. Lochkovian (Early Devonian). [16] U.K. [16] A climatiid with a massive first dorsal fin spine. [16] Parexus sp.png
Paucicanthus [81] Valid. Lochkovian (Early Devonian). [81] Canada. [81] A MOTH locality acanthodian lacking spines on the pectoral and pelvic fins. [81]
Pechoralepis [2] Valid. Pridoli to Givetian (late Silurian to Middle Devonian). [2] [82] Russia, [2] Antarctica. [82] An acritolepid ischnacanthiform based on scales. [2] [82]
Peregrinosquama [2] Valid.Lower Emsian (Early Devonian). [2] Russia. [2] An indeterminate acanthodian based on scales. [2]
Persacanthus Valid. Frasnian (Late Devonian). [15] Iran, Russia, North America. [15] An ischnacanthid very closely related to Atopacanthus . [15]
Pinnacanthus Valid. Emsian (Early Devonian). [19] Wyoming, U.S.A. [19] An acanthodian based on fin spines. [19]
Plectrodus Valid. Ludlow (late Silurian). [30] England. [30] An ischnacanthid commonly confused with Ischnacanthus . [21] [30]
Protodus Valid (non-acanthodian).Late Pragian to Emsian (Early Devonian). [83] Canada. [83] Chondrichthyan teeth originally misidentified as belonging to acanthodiians, due to their similarity to the teeth of a Nostolepis specimen from Scotland. [83] [84]
Podoliacanthus [39] Valid. Pridoli? to Lochkovian (late Silurian? to Early Devonian). [39] Greenland, Ukraine. [39] A small ischnacanthid. [39]
Poracanthodes Valid. Ludlow? to Pragian (late Silurian to Early Devonian). [85] Europe, North America. [85] Namesake of the ischnacanthiform family Poracanthodidae. [85] [3]
Promesacanthus [86] Valid. Lochkovian (Early Devonian). [86] Canada. [86] A mesacanthid acanthodiform from the MOTH locality, with small prepectoral spines similar to "climatiiforms". [86]
Protogonacanthus Valid. Frasnian (Late Devonian). [42] Germany. [42] A cheiracanthid acanthodiform preferring a marine environment. [42]
Pruemolepis Valid? Emsian to Eifelian (Early Devonian to Middle Devonian). [22] [11] Europe. [22] [11] A possible climatiid based on scales similar to some species of Nostolepis . [22]
Pseudacanthodes Valid or junior synonym. Moscovian (Pennsylvanian / Late Carboniferous). [87] Czechia. [87] An eel-like acanthodid acanthodiform, possibly a junior synonym of Traquairichthys . [81] [65] Pseudacanthodes pinnatus.png
Ptomacanthus Valid. Lochkovian (Early Devonian). [88] England, [88] Ukraine. [89] One of the few non- Acanthodes acanthodians with a well-described braincase and gill system, helping to clarify the group's relations with other fishes. [88] [89] Ptomacanthus anglicus.jpg
Ptychodictyon Valid. Emsian to Givetian (Early Devonian to Middle Devonian). [19] Europe, U.S.A. [19] A diplacanthid [10] diplacanthiform based on scales. [19]
Pucapampella Valid (non-acanthodian?). Eifelian to Givetian (Middle Devonian). Bolivia.A chondrichthyan-like braincase sometimes positioned among acanthodians. [50]
Radioporacanthodes Valid. Ludlow to Emsian (late Silurian to Early Devonian). [73] [23] Worldwide. [73] [90] [23] [91] A widespread poracanthodid ischnacanthiform primarily based on scales. [91]
Rhadinacanthus Valid. Emsian to Famennian? (Early Devonian to Late Devonian). [34] [11] [33] Europe, China, Canada. [34] [11] A diplacanthid diplacanthiform similar to, but distinct from, Diplacanthus . [34]
Rockycampacanthus [92] Valid.Late Emsian (Early Devonian). [92] Australia. [92] A distinctive marine ischnacanthid. [92]
Rohonilepis [12] Valid. Ludlow to Pridoli (late Silurian). [12] Lithuania, [12] Latvia.A possible ischnacanthid based on scales. [12]
Sabrinacanthus Valid. Lochkovian (Early Devonian). [30] England. [30] A climatiid with a large shoulder girdle. [21]
Serradentus [93] Valid.Late Givetian (Middle Devonian). [93] Spitsbergen. [93] A medium-sized ischnacanthiform. [93]
Sevyacanthus [19] Valid. Emsian (Early Devonian). [19] U.S.A. [19] A climatiid with complex spine plates. [19]
Sinacanthus Valid (non-acanthodian). Silurian to Early Devonian. Asia. [94] Namesake of the sinacanthids, a group of spined gnathostomes which are likely chondrichthyans. [94]
Striacanthus Valid. Pragian to Frasnian (Early Devonian to Late Devonian). [95] Europe, [95] Australia. [32] A diplacanthiform based on elongated fin spines. [95]
Taemasacanthus [92] Valid.Late Emsian (Early Devonian). [92] Australia. [92] A moderately large marine ischnacanthid. [92]
Taimyrolepis Valid. Pridoli to Lochkovian (late Silurian to Early Devonian). [55] Russia. [55] A possible poracanthodid ischnacanthiform based on scales. [55]
Tareyacanthus Valid. Pragian to Emsian (Early Devonian). [24] [12] Europe, [12] Australia. [24] A possible climatiid based on scales. [24]
Tchunacanthus Valid.Late Llandovery (early Silurian). [2] Russia. [2] A tchunacanthid based on scales. [2]
Teneracanthus [96] Valid.Late Emsian to Famennian (Early Devonian to Late Devonian). [96] [79] Australia, [96] Belgium. [79] A mesacanthid acanthodiform closely related to Lodeacanthus . [96]
Tetanopsyrus [97] Valid. Lochkovian (Early Devonian). [97] Canada. [97] A distinctive diplacanthiform from the MOTH locality. [97] Tetanopsyrus lindoei.png
Traquairia Junior synonym. Moscovian (Pennsylvanian / Late Carboniferous). [87] Czechia.Junior synonym of Traquiairichthys . [98]
Traquairichthys Valid. Moscovian to Leonardian? (Pennsylvanian / Late Carboniferous to Early Permian). [87] [98] Czechia, [87] Greenland?, Texas, U.S.A.? [98] An eel-like acanthodid acanthodiform. [65]
Triazeugacanthus Valid. Frasnian (Late Devonian). [99] Canada. [99] A mesacanthid acanthodiform from the Miguasha Lagerstätte, with one of the best-known ontogenetic series of all acanthodians. [99]
Tricuspicanthus [41] Valid. Lochkovian (Early Devonian). [41] Canada. [41] A small MOTH locality ischnacanthid. [41]
Trundlelepis [24] Valid. Lochkovian to Pragian (Early Devonian). [24] Australia, [24] Nevada, U.S.A.? [90] A poracanthodid ischnacanthiform based on scales. [39]
Uraniacanthus Valid. Lochkovian (Early Devonian). [51] U.K., Canada? [51] A long-spined gladiobranchid diplacanthiform, sometimes misidentified as an ischnacanthiform. [51]
Utahacanthus [100] Valid. Namurian (early Pennsylvanian / Late Carboniferous). [100] Utah, U.S.A. [100] An acanthodid acanthodiform with three prominent otoliths. [100]
Vernicomacanthus Valid. Lochkovian (Early Devonian). [30] U.K. [30] A rare climatiid with denticulated fin spines. [16]
Vesperalia [12] Valid.Late Pridoli (late Silurian). [13] Lithuania. [12] A vesperaliid based on scales. [2]
Watsonacanthus Valid. Emsian to Eifelian (Early Devonian to Middle Devonian). [1] Europe, [1] Australia. [101] A possible climatiid based on scales. [1]
Westrichus [102] Valid.Carboniferous - Permian boundary Germany An acanthodiform. Westrichus kraetschmeri.jpg
Wetteldorfia Valid. Lochkovian to Eifelian (Early Devonian to Middle Devonian). [22] Germany, Spain, Australia, Saudi Arabia. [22] A possible climatiid based on scales. [22]
Xylacanthus Valid.Late Wenlock (mid-Silurian) to Pragian (Early Devonian). [59] Spitsbergen, Canada. [59] An ischnacanthid containing both the largest species of acanthodians (X. grandis at 2.5 meters long) and one of the oldest species (X. kenstewarti). [3] [59]
Yealepis [103] Valid. Ludlow (late Silurian). [103] Australia. [103] A large but indeterminate acanthodian which seems to lack fin spines. [103] [81]
Zemlyacanthus Valid. Lochkovian (Early Devonian). [67] Severnaya Zemlya, Russia. [3] One of the few poracanthodid ischnacanthiforms known from articulated skeletons. [3]


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Chondrichthyes is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or bony fish, which have skeletons primarily composed of bone tissue. Chondrichthyes are aquatic vertebrates with paired fins, paired nares, placoid scales, conus arteriosus in the heart, and a lack of opercula and swim bladders. Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates.

<span class="mw-page-title-main">Gnathostomata</span> Infraphylum of vertebrates

Gnathostomata are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. In addition to opposing jaws, living gnathostomes have true teeth, paired appendages, the elastomeric protein of elastin, and a horizontal semicircular canal of the inner ear, along with physiological and cellular anatomical characters such as the myelin sheaths of neurons, and an adaptive immune system that has the discrete lymphoid organs of spleen and thymus, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene.

<span class="mw-page-title-main">Acanthodii</span> Class of fishes (fossil)

Acanthodii or acanthodians is an extinct class of gnathostomes. They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans and chondrichthyans. In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians.

<i>Climatius</i> Extinct genus of cartilaginous fishes

Climatius is an extinct genus of spiny shark. This genus is known from the Early Devonian (Lochkovian) of Europe, previously considered Silurian remains actually belong to Nostolepis instead.

The Climatiiformes is an order of extinct fish belonging to the class Acanthodii. Like most other "spiny sharks", the Climatiiformes had sharp spines. These animals were often fairly small in size and lived from the Late Silurian to the Early Carboniferous period. The type genus is Climatius. The order used to be subdivided into the suborders Climatiida and Diplacanthida, but subsequently Diplacanthida has been elevated to a separate order, the Diplacanthiformes. The Diplacanthiformes take their name from Diplacanthus, first described by Agassiz in 1843. Family Gyracanthidae is sometimes rejected from this order.

<i>Psarolepis</i> Extinct genus of fishes

Psarolepis is a genus of extinct bony fish which lived around 397 to 418 million years ago. Fossils of Psarolepis have been found mainly in South China and described by paleontologist Xiaobo Yu in 1998. It is not known certainly in which group Psarolepis belongs, but paleontologists agree that it probably is a basal genus and seems to be close to the common ancestor of lobe-finned and ray-finned fishes. In 2001, paleontologist John A. Long compared Psarolepis with onychodontiform fishes and refer to their relationships.

The Xitun Formation is a palaeontological formation which is named after Xitun village in Qujing, a location in South China. This formation includes many remains of fossilized fish and plants of the Early Devonian period. It was originally referred to as the Xitun Member of the Cuifengshan Formation.

<i>Barameda</i> Extinct genus of tetrapodomorphs

Barameda is a genus of rhizodont lobe-finned fishes which lived during the Tournaisian stage near the start of the Carboniferous period in Australia; fossils of the genus have been reported from the Snowy Plains Formation. The largest member of this genus, Barameda decipiens, reached an estimated length of around 3–4 metres (9.8–13.1 ft), while smallest species, B. mitchelli is estimated to have had a length of about 35 centimetres (14 in).

<span class="mw-page-title-main">Paleontology in Oklahoma</span>

Paleontology in Oklahoma refers to paleontological research occurring within or conducted by people from the U.S. state of Oklahoma. Oklahoma has a rich fossil record spanning all three eras of the Phanerozoic Eon. Oklahoma is the best source of Pennsylvanian fossils in the United States due to having an exceptionally complete geologic record of the epoch. From the Cambrian to the Devonian, all of Oklahoma was covered by a sea that would come to be home to creatures like brachiopods, bryozoans, graptolites and trilobites. During the Carboniferous, an expanse of coastal deltaic swamps formed in areas of the state where early tetrapods would leave behind footprints that would later fossilize. The sea withdrew altogether during the Permian period. Oklahoma was home a variety of insects as well as early amphibians and reptiles. Oklahoma stayed dry for most of the Mesozoic. During the Late Triassic, carnivorous dinosaurs left behind footprints that would later fossilize. During the Cretaceous, however, the state was mostly covered by the Western Interior Seaway, which was home to huge ammonites and other marine invertebrates. During the Cenozoic, Oklahoma became home to creatures like bison, camels, creodonts, and horses. During the Ice Age, the state was home to mammoths and mastodons. Local Native Americans are known to have used fossils for medicinal purposes. The Jurassic dinosaur Saurophaganax maximus is the Oklahoma state fossil.

<i>Brochoadmones</i> Extinct genus of cartilaginous fishes

Brochoadmones is an extinct genus of acanthodian from the Devonian of what is now Canada. It is the only genus in the suborder Brochoadmonoidei, whose relationship to other acanthodian orders remains currently in flux.

<i>Tetanopsyrus</i> Extinct genus of cartilaginous fishes

Tetanopsyrus is a genus of Acanthodii. There are two species of this genus that lived through the lower Devonian (Lochkovian), some 416 to 359 million years ago.

The Wood Bay Series is a geologic formation found on the island of Spitsbergen, Svalbard in Norway. It preserves fossils dating back to the Pragian–Emsian stages of the Devonian period.

<i>Gyracanthides</i> Extinct genus of cartilaginous fishes

Gyracanthides is an extinct genus of acanthodian gnathostome, known from Devonian to Early Carboniferous.

<i>Diplacanthus</i> Extinct genus of Devonian spiny sharks.

Diplacanthus is an extinct genus of Mid to Late Devonian fish in the class Acanthodii, known as spiny sharks.

<span class="mw-page-title-main">Diplacanthiformes</span> Extinct order of cartilaginous fishes

Diplacanthiformes is an order of "acanthodian" stem-chondrichthyans which lived during the Devonian Period.

<span class="mw-page-title-main">Acanthodiformes</span> Extinct order of fishes

Acanthodiformes is an order of acanthodian fishes which lived from the Early Devonian to Early Permian.

<span class="mw-page-title-main">Gyracanthidae</span> Extinct family of acanthodian

Gyracanthidae is an family of extinct fish belonging to the class Acanthodii, known from early Devonian to late Carboniferous. Members are characterized by large, broad-based, paired fin spines with the pectoral fin spines having a distinct longitudinal curvature. Although it is originally classified in order Climatiiformes, but later study questioned this.

<i>Qianodus</i> Extinct Silurian chondrichthyan genus

Qianodus is a jawed vertebrate genus that is based on disarticulated teeth from the lower Silurian of China. The type and only species of Qianodus, Q. duplicis, is known from compound dental elements called tooth whorls, each consisting of multiple tooth generations carried by a spiral-shaped base. The tooth whorls of Qianodus represent the oldest unequivocal remains of a toothed vertebrate, predating previously recorded occurrences by about 14 million years. The specimens attributed to the genus come from limestone conglomerate beds of the Rongxi Formation exposed near the village of Leijiatun, Guizhou Province, China. These horizons have been interpreted as tidal deposits1 that form part of the shallow marine sequences of the Rongxi Formation.

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