Fanjingshania Temporal range: Llandovery, | |
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Fossil specimen, pectoral girdle | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | † Acanthodii |
Order: | † Climatiiformes |
Genus: | † Fanjingshania Plamen S. Andreev, 2022 |
Species: | †F. renovata |
Binomial name | |
†Fanjingshania renovata Plamen S. Andreev, 2022 | |
Fanjingshania is an extinct genus of "acanthodian" stem-chondrichthyans from the lower Silurian (late Aeronian, Llandovery Epoch) of China around 439 million years old, making it currently the oldest known acanthodian. It comprises a single species, Fanjingshania renovata which is known from over 1,000 isolated elements, including fin spines, branchiostegal plates, sclerotic plates, trunk scales, and tectal and postorbital tesserae. No teeth were ascribed to the taxon, but in the same horizon the tooth taxon Qianodus was found, and these taxa may be synonymous. In a phylogenetic analysis accompanying the description, it was placed in a clade with Climatius, Parexus, and Ptomacanthus. [1]
Chondrichthyes is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or bony fish, which have skeletons primarily composed of bone tissue. Chondrichthyes are aquatic vertebrates with paired fins, paired nares, placoid scales, conus arteriosus in the heart, and a lack of opercula and swim bladders. Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates.
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Gnathostomata are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. Most gnathostomes have retained ancestral traits like true teeth, a stomach, and paired appendages. Other traits are elastin, a horizontal semicircular canal of the inner ear, myelin sheaths of neurons, and an adaptive immune system which has discrete lymphoid organs, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene.
Acanthodii or acanthodians is an extinct class of gnathostomes. They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans and chondrichthyans. In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians.
Placoderms are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates, and the rest of the body was scaled or naked depending on the species.
Teleostomi is an obsolete taxon of jawed vertebrates that supposedly includes the tetrapods, bony fish, and the wholly extinct acanthodian fish. Key characters of this group include an operculum and a single pair of respiratory openings, features which were lost or modified in some later representatives. The teleostomes include all jawed vertebrates except the chondrichthyans and the extinct class Placodermi.
Galeaspida is an extinct taxon of jawless marine and freshwater fish. The name is derived from galea, the Latin word for helmet, and refers to their massive bone shield on the head. Galeaspida lived in shallow, fresh water and marine environments during the Silurian and Devonian times in what is now Southern China, Tibet and Vietnam. Superficially, their morphology appears more similar to that of Heterostraci than Osteostraci, there being currently no evidence that the galeaspids had paired fins. A galeaspid Tujiaaspis vividus from the Silurian period of China was described in 2022 as having a precursor condition to the form of paired fins seen in Osteostraci and gnathostomes. Earlier than this, Galeaspida were already in fact regarded as being more closely related to Osteostraci, based on the closer similarity of the morphology of the braincase.
Stethacanthus is an extinct genus of shark-like cartilaginous fish which lived from the Late Devonian to Late Carboniferous epoch, dying out around 298.9 million years ago. Fossils have been found in Australia, Asia, Europe and North America.
Acanthodes is an extinct genus of acanthodian fish. Species have been found in Europe, North America, and Asia, spanning the Early Carboniferous to the Early Permian, making it one of the youngest known acanthodian genera.
Psarolepis is a genus of extinct bony fish which lived around 397 to 418 million years ago. Fossils of Psarolepis have been found mainly in South China and described by paleontologist Xiaobo Yu in 1998. It is not known certainly in which group Psarolepis belongs, but paleontologists agree that it probably is a basal genus and seems to be close to the common ancestor of lobe-finned and ray-finned fishes. In 2001, paleontologist John A. Long compared Psarolepis with onychodontiform fishes and refer to their relationships.
The Fanjingshan or Mount Fanjing, located in Tongren, Guizhou province, is the highest peak of the Wuling Mountains in southeastern China, at an elevation of 2,570 m (8,430 ft). The Fanjingshan National Nature Reserve was established in 1978 and designated a UNESCO Biosphere Reserve in 1986. Fanjingshan is a sacred mountain in Chinese Buddhism, considered to be the bodhimaṇḍa of the Maitreya Buddha. It became a UNESCO World Heritage Site in 2018.
Guiyu oneiros is one of the earliest articulated bony fish discovered. Fossils of Guiyu have been found in what is now Qujing, Yunnan, China, in late Silurian marine strata, about 425 million years old.
The Xiushan Formation is a palaeontological formation located in China. It dates to the Lower Silurian period. Studies of conodonts found in the formation suggests the lower member and main part of the upper member correspond to the Pterospathodus eopennatus Superbiozone, whereas the top of the upper member likely correlates with the Pterospathodus celloni Superbiozone.
Brochoadmones is an extinct genus of acanthodian from the Devonian of what is now Canada. It is the only genus in the suborder Brochoadmonoidei, whose relationship to other acanthodian orders remains currently in flux.
Ctenacanthus is an extinct genus of ctenacanthiform chondrichthyan. Remains have been found in the Bloyd Formation of Arkansas and the Cleveland Shale of Ohio in the United States and in South America.
Gyracanthides is an extinct genus of acanthodian gnathostome, known from Devonian to Early Carboniferous.
Shimenolepis granifera is an extinct yunnanolepid placoderm from the Xiaoxi Formation, Li County, Hunan, China. Its age is discussed, while originally considered as late Llandovery, it is later considered to belong to the Ludlow Epoch instead. It was the first described Silurian placoderm, and was the earliest known placoderm until Xiushanosteus was described, known from distinctively ordered plates.
This list of fossil fish research presented in 2022 is a list of new fossil taxa of jawless vertebrates, placoderms, cartilaginous fishes, bony fishes, and other fishes that were described during the year, as well as other significant discoveries and events related to paleoichthyology that occurred in 2022.
Qianodus is a jawed vertebrate genus that is based on disarticulated teeth from the lower Silurian of China. The type and only species of Qianodus, Q. duplicis, is known from compound dental elements called tooth whorls, each consisting of multiple tooth generations carried by a spiral-shaped base. The tooth whorls of Qianodus represent the oldest unequivocal remains of a toothed vertebrate, predating previously recorded occurrences by about 14 million years. The specimens attributed to the genus come from limestone conglomerate beds of the Rongxi Formation exposed near the village of Leijiatun, Guizhou Province, China. These horizons have been interpreted as tidal deposits1 that form part of the shallow marine sequences of the Rongxi Formation.