The Carboniferous is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period 358.9 Ma to the beginning of the Permian Period, 298.9 Ma. It is the fifth and penultimate period of the Paleozoic and the fifth period of the Phanerozoic. In North America, the Carboniferous is often treated as two separate geological periods, the earlier Mississippian and the later Pennsylvanian.
Tullimonstrum, colloquially known as the Tully monster or sometimes Tully's monster, is an extinct genus of soft-bodied bilaterian animal that lived in shallow tropical coastal waters of muddy estuaries during the Pennsylvanian geological period, about 300 million years ago. A single species, T. gregarium, is known. Examples of Tullimonstrum have been found only in the Essex biota, a smaller section of the Mazon Creek fossil beds of Illinois, United States. Its classification has been the subject of controversy, and interpretations of the fossil have likened it to molluscs, arthropods, conodonts, worms, tunicates, and vertebrates. This creature had a mostly cigar-shaped body, with a triangular tail fin, two long stalked eyes, and a proboscis tipped with a mouth-like appendage. Based on the fossils, it seems this creature was a nektonic carnivore that hunted in the ocean’s water column. When Tullimonstrum was alive, Illinois was a mixture of ecosystems like muddy estuaries, marine environments, and rivers and lakes. Fossils of other organisms like crustacean Belotelson, the cnidarian Essexella, and the elasmobranch fish Bandringa have been found alongside Tullimonstrum.
Chimaeras are cartilaginous fish in the order Chimaeriformes, known informally as ghost sharks, rat fish, spookfish, or rabbit fish; the last three names are not to be confused with rattails, Opisthoproctidae, or Siganidae, respectively.
The Ganges shark is a critically endangered species of requiem shark found in the Ganges River and the Brahmaputra River of India and Bangladesh. It is often confused with the more common bull shark, which also inhabits the Ganges River and is sometimes incorrectly referred to as the Ganges shark. The genus is currently considered to contain three recent species; genetic evidence has shown that both the Borneo river shark and Irrawaddy river shark should be regarded as synonyms of the Ganges shark, expanding the range of the species to Pakistan, Myanmar, Borneo, and Java. While the other members of the genus Glyphis occur in coastal marine waters as well as rivers, the Ganges shark is found only in fresh water, making it the world's only exclusively freshwater shark. The species remains poorly known and very rare.
The Mazon Creek fossil beds are a conservation lagerstätte found near Morris, in Grundy County, Illinois. The fossils are preserved in ironstone concretions, formed approximately 309 million years ago in the mid-Pennsylvanian epoch of the Carboniferous period. These concretions frequently preserve both hard and soft tissues of animal and plant materials, as well as many soft-bodied organisms that do not normally fossilize. The quality, quantity and diversity of fossils in the area, known since the mid-nineteenth century, make the Mazon Creek lagerstätte important to paleontologists attempting to reconstruct the paleoecology of the sites. The locality was declared a National Historic Landmark in 1997.
Cobelodus is an extinct genus of cartilaginous fish known from the late Carboniferous to the early Permian period. The type specimen, assigned to the genus Styptobasis, was discovered by Edward Drinker Cope in Illinois Basin black coal shales. Rainer Zangerl reassigned S. aculeata in 1973 to the genus Cobelodus, translating to 'needle tooth'. Cope's description was based from a tooth fragment and was compared to the genus Monocladodus. Cobelodus differs from Styptobasis and Monocladodus in the anatomy of its teeth and pectoral fins.
Pohlsepia mazonensis is a species of fossil organism with unknown affinity. Although it was originally identified as an extinct cephalopod, later studies denied that interpretation. The species is known from a single exceptionally preserved fossil discovered in the late Carboniferous (Pennsylvanian) Francis Creek Shale of the Carbondale Formation, north-east Illinois, United States.
Stethacanthidae is an extinct family of prehistoric holocephalians. It is estimated to have existed approximately between 380 and 300 million years ago. Members of this family are noted for their peculiar dorsal fin.
Iniopteryx is an extinct genus of cartilaginous fish. It is from the Pennsylvanian epoch of the Carboniferous period, approximately 300 million years ago. Their fossils have been found in North America, primarily in two states: Ohio and Montana. In general, very little is known about this species.
Iniopterygiformes is an extinct order of chimaera-like cartilaginous fish that lived from the Devonian to Carboniferous periods. Fossils of them have been found in Montana, Indiana, Illinois, and Nebraska. The Iniopterygians are characterized by large pectoral fins, wing-like projections on their backs, mounted high on the body and denticulated bony plates on the head and jaws. Iniopterygian sharks were small, and their average length was about 18 inches (46 cm). The elongated pectoral fins had denticles along the leading edge which may have had a role in mating. They are thought to have been able to move their pectorals in a vertical plane, ”flying” through the water much like modern-day flying fish.
Cornuboniscus is an extinct genus of prehistoric freshwater ray-finned fish that lived during the Pennsylvanian epoch (Carboniferous), and the only member of the family Cornuboniscidae. It contains a single species, C. budensis from the Bashkirian/lower Westphalian age of what is now Cornwall, England. The genus Cornubonisus was named after the island of Cornubian, and the species name refers to the coastal town of Bude in Cornwall. The type specimen is held in the town's Castle Heritage Centre.
The fossilized remains of fish eggs have an extensive record going at least as far back as the Devonian and spanning into the Cenozoic era. The eggs of many different fish taxa have contributed to this record, including lobe-finned fish, placoderms, and sharks. Occasionally eggs are preserved still within the mother's body, or associated with fossil embryos. Some fossil eggs possibly laid by fish cannot be confidently distinguished from those laid by amphibians; for example, the ichnogenus Mazonova is known from impressions of eggs which resemble eggs of both fish and amphibians. Paleontologist B.K. Hall has observed that the discovery of fossil fish eggs, embryos and larvae link the sciences of paleontology with evo-devo.
Serenichthys kowiensis is a fossil species of coelacanth described in 2015 from near Grahamstown in South Africa.
Ornithoprion is a genus of extinct cartilaginous fish in the family Caseodontidae. The only species, O. hertwigi, lived during the Moscovian stage of the Pennsylvanian, between 315.2 to 307 million years ago, and is known from black shale deposits in what is now the Midwestern United States. The discovery and description of Ornithoprion, performed primarily via radiography, helped clarify the skull anatomy of eugeneodonts; a group which includes O. hertwigi and which were previously known primarily from isolated teeth. The genus derives its name from its pointed, bill-like armored skull and large eyes, which vaguely resemble the features of a bird, while the species name honors Oscar Hertwig. It is known from multiple specimens preserving the skull and frontmost portion of the body.
Pipiscius is an extinct genus of lamprey that lived about 300 million years ago, during the Middle Pennsylvanian Epoch of the Carboniferous Period. The genus contains a single species, P. zangerli, known from the Mazon Creek fossil beds located in present-day Illinois.
Dracopristis is an extinct genus of ctenacanth that lived around 307 million years ago during the Pennsylvanian subperiod of the Carboniferous period. A single species is known, Dracopristis hoffmanorum, which is named in honor of Ralph and Jeanette Hoffman. D. hoffmanorum possessed large dermal denticles along its head, along with rows of short, multi-cusped teeth in its jaws and very large spines on its dorsal fins, features which inspired the etymology of the genus name. The dorsal spines of the holotype fossil are about 0.57 m long, and the whole body was around 2 m long. Its large spines were likely used for defense against larger fishes, while Dracopristis itself was likely a benthic predator which inhabited shallow, brackish-water environments. Before receiving a scientific name, Dracopristis hoffmanorum was informally referred to as the "Godzilla shark".
Romerodus is an extinct genus of cartilaginous fish in the family Caseodontidae. While it and the rest of its family were historically considered elasmobranchs related to sharks and rays, they are now regarded as holocephalans, a diverse subclass which is today only represented by chimaeras. Romerodus is known from the Carboniferous and possibly Permian periods of North America, and the only named species, R. orodontus, was discovered in organic shale deposits in the U.S. state of Nebraska. It is one of few members of its order, the Eugeneodontida, that is known from multiple complete, well preserved body fossils, and is thus an important taxon for understanding the anatomy and ecology of less well preserved eugeneodonts such as Helicoprion. Unlike its larger relatives, the largest known Romerodus were approximately 50 cm (20 in) in total length.
Tanyrhinichthys is an extinct genus of prehistoric ray-finned fish with a lengthened rostrum that lived during the Kasimovian (Missourian) age in what is now New Mexico, United States. Fossils were recovered from Tinajas Member of the Atrasado Formation.
Mayomyzon pieckoensis is an extinct species of lamprey that lived during the Late Carboniferous period, about 300 million years ago. It is the only known species of the genus Mayomyzon, which belongs to the family Mayomyzontidae. It is known from the Mazon Creek fossil beds located in present-day Illinois.
Tyrannophontes is an extinct genus of mantis shrimp that lived during the late Carboniferous period in what is now the Mazon Creek fossil beds of Illinois. It is the only genus in the family Tyrannophontidae. The type species, T. theridion, was described in 1969 by Frederick Schram. A second, much larger species, T. gigantion, was also named by Schram in 2007. Two other species were formerly assigned to the genus, but have since been reclassified.
