Elasmobranchii is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks, rays, skates, and sawfish. Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil.
Stethacanthus is an extinct genus of shark-like holocephalians which lived from the Late Devonian to Late Carboniferous epoch, dying out around 298.9 million years ago. Fossils have been found in Australia, Asia, Europe and North America.
Hybodus is an extinct genus of hybodont, a group of shark-like euselachians that lived from the Late Devonian to the end of the Cretaceous. Species closely related to the type species Hybodus reticulatus lived during the Early Jurassic epoch. Numerous species have been assigned to Hybodus spanning a large period of time, and it is currently considered a wastebasket taxon that is 'broadly polyphyletic' and requires reexamination.
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like cartilaginous fish (chondrichthyans) which existed from the late Devonian to the Late Cretaceous. Hybodonts share a close common ancestry with modern sharks and rays (Neoselachii) as part of the clade Euselachii. They are distinguished from other chondrichthyans by their distinctive fin spines and cephalic spines present on the heads of males. An ecologically diverse group, they were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct.
Lissodus is an extinct genus of hybodont. While fossils attributed to this genus are known spanning from the latest Devonian (Famennian) to the very end of the Cretaceous (Maastrichtian), others consider the genus to have a more narrow range, spanning from the Early Triassic to the end of the Early Cretaceous (Albian). Lissodus is often placed in the family Lonchidiidae, though other authors consider it incertae sedis within Hybodontiformes. Lonchidion has often been regarded as synonymous, but many recent authors consider it to be a distinct but closely related genus. Preserved skull and postcranial remains are known for the Early Triassic species L. cassangensis and L. africanus from Angola and South Africa respectively. Lissodus sensu lato has been estimated to have had a body size range of 15–50 centimetres (0.49–1.64 ft). Remains of Lissodus sensu lato have been found worldwide in both marine and freshwater environments. The low crowned teeth suggest that Lissodus was a bottom dweller that fed by crushing hard shelled organisms (durophagy).
Lonchidion is a genus of extinct hybodont in the family Lonchidiidae. The genus first appears in the fossil record during the Middle Triassic (Ladinian) and was among the last surviving hybodont genera, with its youngest known fossils dating to the very end of the Cretaceous (Maastrichtian).
Meristodonoides is an extinct genus of hybodont. The type species is M. rajkovichi, which was originally a species in the genus Hybodus. The species, along with other Hybodus species such as H. butleri and H. montanensis, was reassigned to Meristodonoides by Charlie J. Underwood and Stephen L. Cumbaa in 2010. The species is primarily known from remains from the Cretaceous of North America, spanning from the Aptian/Albian to Maastrichtian, making it one of the last surviving hybodont genera, though records of the genus likely extend back as far as the Late Jurassic, based on an undescribed skeleton from the Tithonian of England, and fragmentary teeth from the Kimmeridgian of Poland, England and Switzerland. Other remains of the genus are known from the Coniacian of England, and the Aptian-Albian of France. The morphology of the teeth suggests an adaptation to tearing prey. Fossils from the Western Interior Seaway suggest that it preferred nearshore marine environments, being absent from deeper-water areas, with it likely also being able to tolerate brackish and freshwater conditions.
The Eyam Limestone is a geologic formation in the Peak District, England. It preserves fossils dating back to the Viséan stage of the Carboniferous period, and represents a marine environment.
Synechodontiformes is an extinct order of prehistoric shark-like neoselachians, known from the Permian to the Paleogene. They are considered to be members of Neoselachii, the group that contains modern sharks and rays.
Cretodus is an extinct genus of large mackerel sharks belonging to the proposed family Pseudoscapanorhynchidae. Cretodus lived during the Late Cretaceous, ranging from the Cenomanian to the Coniacian. The genus is well-known from strata deposited in the Western Interior Seaway, and from the Late Cretaceous of Europe, Africa, and possibly Asia. Cretodus is primarily represented in the fossil record by isolated teeth and vertebral centra, though a couple of associated dentitions and vertebral columns have been found.
Acanthorhachis is an enigmatic extinct genus of chondrichthyan from the Carboniferous period. Its name is derived from the Greek word acanthos meaning "spine" and the Greek suffix for spine, -rhachis. This is due to the spine-like dermal denticles and their subsidiary spines, which coated the exterior of the animal. The authors who erected this genus suggested the common name "The Spiny Spined Shark." The type species Listracanthus spinatus was in 1896 named by Herbert Bolton.
Listracanthidae is a proposed family of extinct cartilaginous fish. It currently includes the genera Listracanthus and Acanthorhachis. This clade is likely included within Elasmobranchii, but its placement within it is uncertain. Both genera are known from their distinctive spiny dermal dentictles which coat the exterior of their long, slender bodies. They are known from the Viséan to the Early Triassic, mainly from what is now the northern hemisphere. However, there is a possible occurrence in Australia.
Acronemus is an extinct genus of euselachian from the Middle Triassic of Switzerland. It is an enigmatic genus with uncertain relations to other groups. Though originally placed within Ctenacanthiformes, it is now considered Euselachii incertae sedis, due to its mixture of features similar to hybodontiforms and neoselachians. Originally, teeth from this genus were attributed to "Acrodus bicarinatus" while fin spines were named "Nemacanthus tuberculatus". Associated material showed they were the same animal, with the older specific epithet (tuberculatus) taking precedence. The euselachian was given the new genus Acrocnemus, containing a single species. Acronemus is found in the Anisian-age Grenzbitumenzone of Monte San Giorgio. It was a small animal measuring 30–35 cm (0.98–1.15 ft) long. A 2018 study considered it to be closely related to the early Carboniferous genus Tristychius as part of the family Tristychiidae as a basal euselachian, with hybodonts more more closely related to neoselachians than to Tristychiidae.
Amelacanthus is an extinct genus of elasmobranchian cartilaginous fish from the Paleozoic era. It is known from fin spines and currently contains four described species. It is known from the Permian and Carboniferous of North America, Europe, and Africa. It is also known from the Famennian of Russia. Possible specimens of Amelacanthus were found in Permian (Wordian) sediments of Oman.
Cretacladoides is a genus of chondrichthyan, possibly a falcatid, found in France and Austria. Known solely from teeth, mainly found in the Klausrieglerbach locality of Austria, it consists of two species, C. ogiveformis and C. noricum. Assuming a falcatid identity, it is the most recent member of the family, which otherwise became extinct at the end of the Carboniferous.
Squatinactis is a genus of extinct elasmobranch chondrichthyan known from the Carboniferous aged Bear Gulch Limestone in Montana. This fish was discovered in 1974 by Richard Lund. The type specimen, named CMNH 46133, consists of a brain case, poorly preserved jaws and gills, a pectoral fin, and a partial vertebral axis. This creatures most startling feature were its broad pectoral fins which resembled those of stingrays and angel sharks (Squatina). The holotype specimen has about 15 teeth in its jaw. This creature is named after the angel shark. Remains found in the South Urals of Russia and the Eyam Limestone of Derbyshire, England, have been tentatively identified as those belonging to S. caudispinatus.
Egertonodus is an extinct genus of shark-like hybodont fish. It includes E. basanus from the Jurassic of Europe and North Africa and Cretaceous of North America, North Africa and Europe, and E. duffini from the Middle Jurassic of England. Indeterminate remains of the genus have been reported from the Early Cretaceous of Asia. E. basanus is known from preserved skull material, while E. duffini is only known from teeth. The genus is distinguished from Hybodussensu stricto by characters of the skull and teeth. E. basanus, the most common species, is thought to have reached 1.5 m in length. E. fraasi from the Late Jurassic of Germany, known from a poorly preserved full body fossil, was placed in Egertonodus in one study, but this has been subsequently questioned by other authors, due to strong differences in tooth morphology from the type species. Fossils have been found in freshwater and lagoonal environments.
Cooleyella is an extinct genus of cartilaginous fish which lived from the Carboniferous to the Permian of the United States, Russia, UK and Oman. Several species have been described. It is placed in the family Anachronistidae.
Planohybodus is an extinct genus of hybodont, known from the Middle Jurassic-Early Cretaceous (Bathonian-Barremian) of Europe and the Indian subcontinent. Fossils have been found in marine as well as freshwater environments. The genus contains 3 confirmed species, two of which were originally assigned to the genus Hybodus. Possible records have been reported from the Late Jurassic of Mexico, the Early Cretaceous of Brazil and the Late Cretaceous (Santonian) of North America, but these are unconfirmed. Planohybodus peterboroughensis is suggested to have reached lengths of 2–3 metres (6.6–9.8 ft). A specimen of the ammonite genus Orthaspidoceras from the Late Jurassic of France has been found with an embedded tooth of Planohybodus, suggesting that while the teeth of Planohybodus were adapted to tearing soft bodied prey, it would attack hard-shelled prey at least on occasion.
Parvodus is an extinct genus of hybodont, known from the Mesozoic era.
