The Permian is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period 298.9 million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia.
The Uralian orogeny refers to the long series of linear deformation and mountain building events that raised the Ural Mountains, starting in the Late Carboniferous and Permian periods of the Palaeozoic Era, c. 323–299 and 299–251 million years ago (Mya) respectively, and ending with the last series of continental collisions in Triassic to early Jurassic times.
The Cisuralian is the first series/epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the western slopes of the Ural Mountains in Russia and Kazakhstan and dates between 298.9 ± 0.15 – 272.3 ± 0.5 Ma.
In the geologic timescale, the Artinskian is an age or stage of the Permian. It is a subdivision of the Cisuralian Epoch or Series. The Artinskian likely lasted between 290.1 and 283.5 million years ago (Ma) according to the most recent revision of the International Commission on Stratigraphy (ICS) in 2022. It was preceded by the Sakmarian and followed by the Kungurian.
Estemmenosuchidae is an extinct family of large, very early herbivorous therapsids that flourished during the Guadalupian period. They are distinguished by horn-like structures, probably for display or agonistic behavior. Apart from the best known genus, Estemmenosuchus, the group is poorly known. To date, their fossils are known only from the Perm region of Russia.
Estemmenosuchus is an extinct genus of large, early omnivorous therapsid. It is believed and interpreted to have lived during the middle part of the Middle Permian around 267 million years ago. The two species, E. uralensis and E. mirabilis, are characterised by distinctive horn-like structures, which were probably used for intra-specific display. Both species of Estemmenosuchus are from the Perm region of Russia. Two other estemmenosuchids, Anoplosuchus and Zopherosuchus, are now considered females of the species E. uralensis. There were many complete and incomplete skeletons found together.
Rhopalodon is an extinct genus of therapsids from the Permian of Russia. It has been variously classified as a dinosaur, a dinocephalian, or another branch of amniotes. Rhopalodon is notable for being among the first "reptiles" mentioned in Nature. T.H. Huxley wrote of this animal, among others, in the inaugural issue of the magazine, in November 1869. He gave the age of this animal and of the contemporary Deuterosaurus as Triassic, but both are now known to have lived during the Middle Permian.
Archegosauridae is a family of relatively large and long snouted temnospondyls that lived in the Permian period. They were fully aquatic animals, and were metabolically and physiologically more similar to fish than modern amphibians. The family has been divided into two subfamilies, the longer-snouted Platyoposaurinae and the shorter-snouted Melosaurinae.
Rhipaeosaurus is an extinct genus of nycteroleterid parareptile known from an articulated skeleton from the mid Middle Permian of European Russia. It contained a single species, Rhipaeosaurus tricuspidens. A bayesian analysis suggests that it is more closely related to pareiasaurs than to the other nycteroleterids, due to skull and tooth features. For this reason, "Nycteroleteridae" may be a grade rather than a clade, unless redefined to exclude Rhipaeosaurus.
Venyukovia is an extinct genus of venyukovioid therapsid, a basal anomodont from the Middle Permian of Russia. The type and sole species, V. prima, is known only by a partial lower jaw with teeth. Venyukovia has often been incorrectly spelt as 'Venjukovia' in English literature. This stems from a spelling error made by Russian palaeontologist Ivan Efremov in 1940, who mistakenly replaced the 'y' with a 'j', which subsequently permeated through therapsid literature before the mistake was caught and corrected. Venyukovia is the namesake for the Venyukovioidea, a group of small Russian basal anomodonts also including the closely related Otsheria, Suminia, Parasuminia and Ulemica, although it itself is also one of the poorest known. Like other venyukovioids, it had large projecting incisor-like teeth at the front and lacked canines, although the remaining teeth are simple compared to some other venyukovioids, but may resemble those of Otsheria.
Kamacops is a genus of dissorophid temnospondyls known from the Middle to Late Permian of Russia that was described by Yuri Gubin in 1980. It is known from a single species, Kamacops acervalis, material of which is currently reposited in the Paleontological Institute of the Russian Academy of Sciences. Kamacops is one of the youngest dissorophids, along with Iratusaurus and Zygosaurus from Russia and Anakamacops from China and was one of the largest known dissorophids, with an estimated skull length of 24–30 cm. It is typically recovered as being most closely related to Zygosaurus and to the North American Cacops. A detailed study of the braincase region was performed by Schoch (1999), one of the first to examine this region in dissorophids. This study revealed extensive co-ossification of the braincase.
Archegosauroidea is an extinct superfamily of Permian temnospondyls. The superfamily is assigned to the clade Stereospondylomorpha and is the sister taxon to the suborder Stereospondyli. It includes the families Actinodontidae and Archegosauridae, and possibly the genus Intasuchus, which is placed within the monotypic family Intasuchidae. They were fully aquatic animals, and were metabolically and physiologically more similar to fish than modern amphibians.
Northwest Russia, or the Russian North is the northern part of western Russia. It is bounded by Norway, Finland, the Arctic Ocean, the Ural Mountains and the east-flowing part of the Volga. The area is roughly coterminous with the Northwestern Federal District, which it is administered as part of. Historically, it was the area of the Novgorod and Pskov merchant republics.
Idelinellidae is a family of fossil cockroach-like insects. They closely resemble members of the family Stenoneuridae, and consist of the following species:
Austrolimulidae is an extinct family of horseshoe crabs belonging to the infraorder Limulina. Members of the family are known from the Permian to the beginning of the Jurassic, though one species has been reported from the end of the Cretaceous. Austrolimulids are known for amongst the most extreme morphologies among Xiphosurids, including large elongated genal spines. Unlike living Limulids, Austrolimulids were likely adapted for freshwater and brackish environments. They are considered to be the sister group to Limulidae, the group that contains all modern horseshoe crabs.
Prostacheoceras is genus of ammonoid cephalopods belonging to the Vidrioceratidae family. Species belonging to this genus lived in early to middle Permian.
Sweetognathus is an extinct genus of conodonts in the family Sweetognathidae that evolved at the beginning of the Permian period, in near-equatorial, shallow-water seas.
The geology of Kazakhstan includes extensive basement rocks from the Precambrian and widespread Paleozoic rocks, as well as sediments formed in rift basins during the Mesozoic.
Adamantina is an extinct genus of jalodont chondrichthyan from the Carboniferous and Permian periods. It is known mainly from isolated teeth and scales. It contains two species, A. foliacea and A. benedictae. The type species, A. benedictae, is known from the Wuchiapingian of east Greenland and the Roadian of the Kanin Peninsula, Russia. A. foliacea is known from the late Tournaisian, late Artinskian, and the early Asselian of Russia and the late Kasimovian of Iowa, United States. There is also a likely occurrence in the upper Pennsylvanian of Brazil.
Bransonella is an extinct genus of marine Xenacanth which lived during the Paleozoic era. It is known only from teeth which are easily distinguished from related genera by ornamentation on the cusp shaped like an inverted "V" and fin spines Teeth attributed to this genus are small, no greater than 2 millimeters. This suggests a small body size likely not exceeding a meter in length. It has been suggested their lifestyle was similar to modern Catsharks. It was used to erect a new order along with Barbclabornia based on ornamentation.
