Carinacanthus Temporal range: | |
---|---|
Type specimen | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Chondrichthyes |
Subclass: | Elasmobranchii |
Order: | † Hybodontiformes |
Genus: | † Carinacanthus Bryant, 1934 |
Species: | †C. jepseni |
Binomial name | |
†Carinacanthus jepseni Bryant, 1934 | |
Carinacanthus ("keeled spine") is an extinct genus of small freshwater hybodont shark that inhabited eastern North America during the Triassic period. [1] [2] It contains a single species, C. jepseni from the Late Triassic (Norian)-aged Lockatong Formation of Pennsylvania, USA. It was named after paleontologist Glenn Lowell Jepsen. [3] Some authors consider it a late-surviving ctenacanthiform. [1] [4] [5]
It is a very rare taxon, with only two potential specimens known despite the well-studied nature of the Newark Supergroup deposits. The type specimen consists of a poorly-preserved axial skeleton, only about 130 millimetres (5.1 in) in length, which is missing the head but allows most of the post-cranial anatomy to be known, including the position of its two large spines and the presence of a heterocercal tail. In addition to this type specimen, a second potential specimen is known, consisting of a poorly-preserved head with only the jaw and teeth recognizable. [3]
Despite the poor preservation of the type specimen, it is one of the only Triassic elasmobranchs from the United States to be known from articulated fossils instead of isolated teeth & spines, and was the first Triassic elasmobranch overall to be described from the eastern United States. It and Palaeobates were the only freshwater hybodonts known from the Triassic of the United States for a significant period of time, although more recent studies have also found the presence of small hybodont teeth in North Carolina. [6] [7]
The fossil egg case taxon Chimaerotheca oakesi , described from the Triassic of Massachusetts and attributed to a chimaeroid, may be potentially referable to Carinacanthus, as one of the only other chondrichthyans known from the Newark Supergroup. [8]
Elasmobranchii is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks, rays, skates, and sawfish. Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil.
Cladoselache is an extinct genus of shark-like chondrichthyan from the Late Devonian (Famennian) of North America. It was similar in body shape to modern lamnid sharks, but was not closely related to lamnids or to any other modern (selachian) shark. As an early chondrichthyan, it had yet to evolve traits of modern sharks such as accelerated tooth replacement, a loose jaw suspension, enameloid teeth, and possibly claspers.
Xenacanthida is an order or superorder of extinct shark-like chondrichthyans known from the Carboniferous to Triassic. They were native to freshwater, marginal marine and shallow marine habitats. Some xenacanths may have grown to lengths of 5 m (16 ft). Most xenacanths died out at the end of the Permian in the End-Permian Mass Extinction, with only a few forms surviving into the Triassic.
Hybodus is an extinct genus of hybodont that lived from the Middle Triassic to the Late Cretaceous periods. Species closely related to the type species Hybodus reticulatus lived during the Early Jurassic epoch. Numerous species have been assigned to Hybodus spanning a large period of time, and it is currently considered a wastebasket taxon that is 'broadly polyphyletic' and requires reexamination.
Falcatus is an extinct genus of falcatid chondrichthyan which lived during the early Carboniferous Period in Bear Gulch bay in what is now Montana.
Diplurus is a genus of prehistoric mawsoniid coelacanth fish which lived during the Late Triassic-Early Jurassic period in North America. The fossils of this genus are common on the eastern North American Margin, being a key taxon of the Newark Supergroup, and recovered from units such as the Bull Run Formation, Lockatong Formation, Stockton Formation, Solite Quarry, Midland Formation, East Berlin Formation, Boonton Formation and the Portland Formation. Three species are know, the type D. longicaudatus, the youngest and biggest, D. newarki, the oldest, followed then by "D. uddeni" (Eastman), also from older rocks and considered dubious. A recent work has recovered a 3rd or 4th species, D. enigmaticus, from the Late Triassic of New Jersey, representing another small-bodied form living in sympatry with the similarly sized D. newarki. This genus ranges in size from 15 cm of D. newarki and D. enigmaticus in New Jersey to the larger 60 cm specimens of Diplurus longicaudatus found in the Connecticut River Valley, which indicates a considerable growth and likely a change in the ecological position of the genus to a possible apex predatory niche.
Gnathorhiza is an extinct genus of prehistoric lobe-finned fish (lungfish) which lived from the Carboniferous period to the Early Triassic epoch. It is the only known lungfish genus to have crossed the Permo-Triassic boundary. Several species have been described, ranging in size from 5 to 50 centimeters.
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like cartilaginous fish (chondrichthyans) which existed from the late Devonian to the Late Cretaceous. Hybodonts share a close common ancestry with modern sharks and rays (Neoselachii) as part of the clade Euselachii. They are distinguished from other chondrichthyans by their distinctive fin spines and cephalic spines present on the heads of males. An ecologically diverse group, they were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct.
The Cow Branch Formation is a Late Triassic geologic formation in Virginia and North Carolina in the eastern United States. The formation consists of cyclical beds of black and grey lacustrine (lake) mudstone and shale. It is a konservat-lagerstätte renowned for its exceptionally preserved insect fossils, along with small reptiles, fish, and plants. Dinosaur tracks have also been reported from the formation.
Lonchidion is a genus of extinct hybodont in the family Lonchidiidae. The genus first appears in the fossil record during the Middle Triassic (Ladinian) and was among the last surviving hybodont genera, with its youngest known fossils dating to the very end of the Cretaceous (Maastrichtian).
Palaeoxyris is a morphogenus of eggs cases, widely thought to have been produced by hybodonts, with a predominant occurrence in ancient freshwater environments. They comprise a beak, a body and a pedicle. They display a conspicuous right-handed spiral of collarettes around the body, and in some cases, the pedicle, resulting in a rhomboidal pattern when flattened during fossilisation. At the end of the beak was a tendril which attached the egg to vegetation during development. The body of the egg ranges in length from 1.2–8.9 centimetres (0.47–3.50 in), depending on the species.
Ornithoprion is a monotypic genus of extinct eugeneodont chondrichthyan in the family Caseodontidae. The type and only species, O. hertwigi, lived during the Moscovian stage of the Pennsylvanian, between 315.2 to 307 million years ago, and is known from black shale deposits in what is now the Midwestern United States. The discovery and description of Ornithoprion, performed primarily via radiography, helped clarify the anatomy of the eugeneodonts, a group which were previously known primarily from isolated tooth whorls. The genus derives its name from its pointed, bill-like armored skull and large eyes, which vaguely resemble the features of a bird, while the species name honors Oscar Hertwig. It is known from specimens preserving the skull and anterior portion of the body.
Synechodontiformes is an extinct order of prehistoric shark-like cartilaginous fish, known from the Permian to the Paleogene. They are considered to be members of Neoselachii, the group that contains modern sharks and rays.
Tribodus is an extinct genus of hybodont. It lived during the mid Cretaceous (Albian-Cenomanian) with fossils being known from northern South America, North Africa, and southern Europe.
Palaeobates is an extinct genus of prehistoric elasmobranchs in the order Hybodontiformes. It lived during the Triassic period. It was a small shark about 1 m (3.3 ft) long. Palaeobates had a grinding-type dentition, which it used to crush hard-shelled prey. The teeth exhibit an orthodont histology.
Arctacanthus is an extinct genus of cartilaginous fish from the Permian and possibly Triassic periods.
Romerodus is an extinct genus of cartilaginous fish in the family Caseodontidae. While it and the rest of its family were historically considered elasmobranchs related to sharks and rays, they are now regarded as holocephalans, a diverse subclass which is today only represented by chimaeras. Romerodus is known from the Carboniferous and possibly Permian periods of North America, and the type and only named species, R. orodontus, was discovered in organic shale deposits in the U.S. state of Nebraska. It is one of few members of its order, the Eugeneodontida, which is known from multiple complete, well preserved body fossils, and is thus an important taxon for understanding the anatomy and ecology of less well preserved eugeneodonts such as Helicoprion. Unlike its larger relatives, Romerodus grew to approximately 50 cm in total length.
Anachronistidae is an extinct family of cartilaginous fish, known from the Carboniferous and Permian periods. They are considered to be the oldest known members of Neoselachii, with a close relationship to modern sharks and rays. They are known from isolated teeth. They first appeared in Europe during the late Mississippian (Viséan), with Cooleyella dispersing into North America and South America during the Late Carboniferous and Permian.
Gansuselache is a genus of extinct elasmobranch in the order Hybodontiformes, comprising one species, Gansuselache tungshengi (monotypy) from Gansu Province, China. It is known from Fangshankou Formation, which previously interpreted as Permian in age, but reinterpreted as the Early Triassic in later study. A tooth remain tentatively assigned to this genus is also known from Late Permian strata of Poland.
Cionichthys is an extinct genus of freshwater ray-finned fish that inhabited southwestern and eastern North America during the Late Triassic period. It was a member of the Redfieldiiformes, an order of fishes widespread throughout freshwater habitats at this time, especially in North America.