Acanthodii or acanthodians is an extinct class of gnathostomes. They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans and chondrichthyans. In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians.
Cladoselache is an extinct genus of shark-like chondrichthyan from the Late Devonian (Famennian) of North America. It was similar in body shape to modern lamnid sharks, but was not closely related to lamnids or to any other modern (selachian) shark. As an early chondrichthyan, it had yet to evolve traits of modern sharks such as accelerated tooth replacement, a loose jaw suspension, enameloid teeth, and possibly claspers.
Temnospondyli or temnospondyls is a diverse ancient order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian and Triassic periods, with fossils being found on every continent. A few species continued into the Jurassic and Early Cretaceous periods, but all had gone extinct by the Late Cretaceous. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are amphibians, many had characteristics such as scales and armour-like bony plates that distinguish them from the modern soft-bodied lissamphibians.
Koolasuchus is an extinct genus of brachyopoid temnospondyl in the family Chigutisauridae. Fossils have been found from Victoria, Australia and date back 125-120 million years ago to Barremian-Aptian stages of the Early Cretaceous. Koolasuchus is the youngest known temnospondyl. It is known from several fragments of the skull and other bones such as vertebrae, ribs, and pectoral elements. The type species Koolasuchus cleelandi was named in 1997. K. cleelandi was adopted as the fossil emblem for the state of Victoria, Australia on 13 January 2022.
Stethacanthus is an extinct genus of shark-like holocephalians which lived from the Late Devonian to Late Carboniferous epoch, dying out around 298.9 million years ago. Fossils have been found in Australia, Asia, Europe and North America.
Xenacanthida is an order or superorder of extinct shark-like chondrichthyans known from the Carboniferous to Triassic. They were native to freshwater, marginal marine and shallow marine habitats. Some xenacanths may have grown to lengths of 5 m (16 ft). Most xenacanths died out at the end of the Permian in the End-Permian Mass Extinction, with only a few forms surviving into the Triassic.
The Bear Gulch Limestone is a limestone-rich geological lens in central Montana, renowned for the quality of its late Mississippian-aged fossils. It is exposed over a number of outcrops northeast of the Big Snowy Mountains, and is often considered a component of the more widespread Heath Formation. The Bear Gulch Limestone reconstructs a diverse, though isolated, marine ecosystem which developed near the end of the Serpukhovian age. It is a lagerstätte, a particular type of rock unit with exceptional fossil preservation of both articulated skeletons and soft tissues. Bear Gulch fossils include a variety of fish, invertebrates, and algae occupying a number of different habitats within a preserved shallow bay.
The Climatiiformes is an order of extinct fish belonging to the class Acanthodii. Like most other "spiny sharks", the Climatiiformes had sharp spines. These animals were often fairly small in size and lived from the Late Silurian to the Early Carboniferous period. The type genus is Climatius. The order used to be subdivided into the suborders Climatiida and Diplacanthida, but subsequently Diplacanthida has been elevated to a separate order, the Diplacanthiformes. The Diplacanthiformes take their name from Diplacanthus, first described by Agassiz in 1843. Family Gyracanthidae is sometimes rejected from this order.
Barameda is a genus of rhizodont lobe-finned fishes which lived during the Tournaisian stage near the start of the Carboniferous period in Australia; fossils of the genus have been reported from the Snowy Plains Formation. The largest member of this genus, Barameda decipiens, reached an estimated length of around 3–4 metres (9.8–13.1 ft), while smallest species, B. mitchelli is estimated to have had a length of about 35 centimetres (14 in).
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like cartilaginous fish (chondrichthyans) which existed from the late Devonian to the Late Cretaceous. Hybodonts share a close common ancestry with modern sharks and rays (Neoselachii) as part of the clade Euselachii. They are distinguished from other chondrichthyans by their distinctive fin spines and cephalic spines present on the heads of males. An ecologically diverse group, they were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct.
Brachyopoidea is a superfamily of temnospondyls that lived during the Mesozoic. It contains the families Brachyopidae and Chigutisauridae. The earliest records of brachyopids are from the Lower Triassic in Australia. The latest-surviving member of the superfamily is the chigutisaurid Koolasuchus from the Early Cretaceous of Australia.
Lydekkerinidae is a family of stereospondyl temnospondyls that lived in the Early Triassic period. During this time period, lydekkerinids were widely distributed, with putative remains reported from Russia, Greenland, India, South Africa, Madagascar, Australia, and Antarctica. In contrast to most other stereospondyls, lydekkerinids were relatively small-bodied. The type genus is Lydekkerina, the namesake of the family and the best-known lydekkerinid.
Acroplous is an extinct genus of dvinosaurian Temnospondyli within the family Eobrachyopidae.
Ossinodus is an extinct genus of stem tetrapod. Fossils have been found from the Ducabrook Formation in Queensland, Australia dating back to the middle Visean stage of the Early Carboniferous (Mississippian). It was originally placed within the family Whatcheeriidae, but the absence of an intertemporal bone as suggested by a recent reconstruction of the skull based on fragmentary material may prove it to be stemward of all whatcheeriids.
Reginaselache is an extinct genus of xenacanthiform known from the Early Carboniferous of central Queensland, Australia. It was found in the middle the Ducabrook Formation, northwest of Springsure. It was first named by Susan Turner and Carole J. Burrow in 2011 and the type species is Reginaselache morrisi.
Thrinacodus is an extinct genus of basal elasmobranch, found worldwide from the Late Devonian-Lower Carboniferous. The type species is Thrinacodus nanus. Most species are only known from their tricuspid teeth. T. gracia, originally placed in the separate genus Thrinacoselache from the Serpukhovian-aged Bear Gulch Limestone, of what is now Montana, is known from full body impressions, showing a long, slender eel-like body up to a metre in length, with an elongate rostrum. Stomach contents of T. gracia include remains of crustaceans and small chondrichthyan fish. It is a member of the Phoebodontiformes.
Gyracanthides is an extinct genus of acanthodian gnathostome, known from Devonian to Early Carboniferous.
Ageleodus is an extinct genus of cartilaginous fish from the Paleozoic era. It is known from two species, both of which are based upon isolated teeth. A. pectinatus is known from the Carboniferous of Europe and North America. A. altus was described in 2006 from the Carboniferous of Australia. Four teeth from the Tournaisian deposits of Canning Basin probably present another Australian species although their shape is likely caused by physical wear rather than species features. This genus is also known from the Famennian of the Catskill formation at the Red Hill Site in Hyner, Pennsylvania. A study of 382 specimens from the site showed the strong heterodonty of this genus, which varies widely in tooth length and cusp number. This study described them as A. pectinatus, but the paper which described A. altus tentatively labeled them Ageleodus cf. A. altus. While generally considered a chondrichthyan, it has eluded classification into any known order or family.
Ctenacanthiformes is an extinct order of elasmobranch fish. They possessed ornamented fin spines at the front of their dorsal fins and cladodont-type dentition, that is typically of a grasping morphology, though some taxa developed cutting and gouging tooth morphologies. Some ctenacanths are thought to have reached sizes comparable to the great white shark, with body lengths of up to 7 metres (23 ft) and weights of 1,500–2,500 kilograms (3,300–5,500 lb). Ctenacanths are typically thought to have existed from the Devonian to the Late Permian, becoming extinct in the Permian-Triassic extinction event. Members of the family Ctenacanthidae may have survived into the Cretaceous based on teeth found in deep water deposits of Valanginian age in France and Austria, however, other authors contend that the similarity of these teeth to Paleozoic ctenacanths is only superficial, and they likely belong to neoselachians instead. The monophyly of the group has been questioned, with some studies recovering the group as a whole as paraphyletic or polyphyletic.
