Chondrichthyes is a class of jawed fish that contains the cartilaginous fish or chondrichthyians, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or bony fish, which have skeletons primarily composed of bone tissue. Chondrichthyes are aquatic vertebrates with paired fins, paired nares, placoid scales, conus arteriosus in the heart, and a lack of opecula and swim bladders. Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates.
Elasmobranchii is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks, rays, skates, and sawfish. Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil.
Helicoprion is an extinct genus of shark-like eugeneodont fish. Almost all fossil specimens are of spirally arranged clusters of the individuals' teeth, called "tooth whorls", which in life were embedded in the lower jaw. As with most extinct cartilaginous fish, the skeleton is mostly unknown. Fossils of Helicoprion are known from a 20 million year timespan during the Permian period from the Artinskian stage of the Cisuralian to the Roadian stage of the Guadalupian. The closest living relatives of Helicoprion are the chimaeras, though their relationship is very distant. The unusual tooth arrangement is thought to have been an adaption for feeding on soft bodied prey, and may have functioned as a deshelling mechanism for hard bodied cephalopods such as nautiloids and ammonoids. In 2013, systematic revision of Helicoprion via morphometric analysis of the tooth whorls found only H. davisii, H. bessonowi and H. ergassaminon to be valid, with some of the larger tooth whorls being outliers.
Xenacanthida is an order or superorder of extinct shark-like chondrichthyans known from the Carboniferous to Triassic. They were native to freshwater, marginal marine and shallow marine habitats. Some xenacanths may have grown to lengths of 5 m (16 ft). Most xenacanths died out at the end of the Permian in the End-Permian Mass Extinction, with only a few forms surviving into the Triassic.
Edestus is an extinct genus of eugeneodontid holocephalian fish known from the Late Carboniferous (Pennsylvanian) of the United Kingdom, Russia, and the United States. Most remains consist of isolated curved blades or "whorls" that are studded with teeth, that in life were situated within the jaws. Edestus is a Greek name derived from the word edeste, in reference to the aberrant quality and size of the species' teeth. The largest species, E. heinrichi, has been conservatively estimated to reach greater than 6.7 m (22 ft) in length, around the size of the largest known great white shark, possibly making it the largest marine predator to have ever existed up to that point.
Sarcoprion is an extinct genus of eugeneodontid holocephalids from the Permian of Greenland. Similar to other eugeneodontids such as Edestus and Helicoprion, it was best known for its extremely bizarre tooth morphology compared to other species of sharks and their closest relatives, the chimaeras. Compared to other members of the Helicoprionidae, its "tooth whorls" were found to be sharper, more compact, and in better condition than other holocephalians of the time, and refrained from growing to extremely unwieldy forms that would raise questions about its ability to feed properly. The genus contains one species, Sarcoprion edax, found in Permian-aged marine strata of Meddelelser om Grønland.
Campodus is an extinct genus of eugeneodont holocephalans from the Carboniferous. Likely one of the earliest and most basal caseodontoids, it can be characterized by its broad, ridge-ornamented crushing teeth made of various types of dentine. The type species, C. agassizianus, was originally described in 1844 based on a small number of teeth from the Namurian of Belgium.
Fadenia is an extinct genus of eugeneodontid holocephalian chondrichthyan from the Carboniferous Period of Missouri, the Permian period of Greenland, and the Early Triassic epoch of British Columbia, Canada.
Caseodus is an extinct genus of eugeneodontid holocephalian from the Carboniferous of the United States and the Early Triassic of Canada. It was of medium size, measuring 1–1.5 m (3.3–4.9 ft) in length.
Parahelicoprion is an extinct genus of shark-like eugeneodontid holocephalids from the Permian of the Ural Mountains (Russia) and Copacabana Formation, Bolivia. The genus name, from "nearly coiled saw" in Greek, directly refers to Helicoprion, a related holocephalid that shares similar traits to it, including the helical whorl of teeth.
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like cartilaginous fish (chondrichthyans) which existed from the late Devonian to the Late Cretaceous. Hybodonts share a close common ancestry with modern sharks and rays (Neoselachii) as part of the clade Euselachii. They are distinguished from other chondrichthyans by their distinctive fin spines and cephalic spines present on the heads of males. An ecologically diverse group, they were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct.
Helicoprionidae is an extinct, poorly known family of bizarre holocephalids within the poorly understood order Eugeneodontida. Members of the Helicoprionidae possessed a unique "tooth-whorl" on the symphysis of the lower jaw and pectoral fins supported by long radials. The closest living relatives of the Helicoprionidae and all other eugeneodontids are the ratfishes. The anatomy of the tooth-whorl differed amongst genus and species, some possessing complete spirals, others possessing halved spirals, and some with wedged half-spirals. Each tooth-whorl is thought to be adapted to a different type of prey, and a different predation strategy.
The Edestidae are a poorly known, extinct family of shark-like eugeneodontid holocephalid cartilaginous fish.
Iniopterygiformes is an extinct order of chimaera-like cartilaginous fish that lived from the Devonian to Carboniferous periods. Fossils of them have been found in Montana, Indiana, Illinois, and Nebraska. The Iniopterygians are characterized by large pectoral fins, wing-like projections on their backs, mounted high on the body and denticulated bony plates on the head and jaws. Iniopterygian sharks were small, and their average length was about 18 inches (46 cm). The elongated pectoral fins had denticles along the leading edge which may have had a role in mating. They are thought to have been able to move their pectorals in a vertical plane, ”flying” through the water much like modern-day flying fish.
This list of fossil fishes described in 2013 is a list of new taxa of placoderms, fossil cartilaginous fishes and bony fishess of every kind that have been described during the year 2013. The list only includes taxa at the level of genus or species.
Mcmurdodus is an extinct genus of chondrichthyan from Antarctica and Australia and the sole member of the family Mcmurdodontidae. It contains two extinct species. However, the Australian species M. whitei has been found to be different from the Antarctic type species M. featherensis, and thus M. whitei has been classified into a new genus, Maiseyodus.
Toxoprion is an extinct genus of eugeneodont holocephalids whose fossils are found in marine strata from the Early Carboniferous until the Late Permian near Eureka, Nevada.
Agassizodus is an extinct genus of eugeneodont holocephalian from the Carboniferous. It belongs to the family Helicoprionidae, which is sometimes called Agassizodontidae. Like other members of its family, it possessed a symphyseal tooth whorl, which was likely present at the tip of the lower jaw and associated with lateral crushing toothplates. The type species, A. variabilis, was originally named Lophodus variabilis until the name "Lophodus" was determined to be preoccupied.
This list of fossil fish described in 2018 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fish, bony fish, and other fish of every kind that are scheduled to be described during the year 2018, as well as other significant discoveries and events related to paleontology of fish that are scheduled to occur in 2018.
Lestrodus is an extinct genus of edestid fish that lived during the Late Carboniferous. It contains one valid species, L. newtoni, which is known from a single tooth whorl from the Millstone Grit of England. It was originally named as a species of Edestus, but is now considered a distinct genus based on morphological differences.
