Synapsids are one of the two major clades of vertebrate animals in the group Amniota, the other being the sauropsids, which include reptiles and birds. The synapsids were the dominant land animals in the late Paleozoic and early Mesozoic, but the only extant group that survived into the Cenozoic are the mammals. Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye orbit, leaving a bony arch beneath each; this accounts for their name. The distinctive temporal fenestra developed about 318 million years ago during the Late Carboniferous period, when synapsids and sauropsids diverged, but was subsequently merged with the orbit in early mammals.
Rhynchocephalia is an order of lizard-like reptiles that includes only one living species, the tuatara of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a speciose group with high morphological and ecological diversity. The oldest record of the group is dated to the Middle Triassic around 238 to 240 million years ago, and they had achieved a worldwide distribution by the Early Jurassic. Most rhynchocephalians belong to the group Sphenodontia ('wedge-teeth'). Their closest living relatives are lizards and snakes in the order Squamata, with the two orders being grouped together in the superorder Lepidosauria.
Helicoprion is an extinct genus of shark-like eugeneodont fish. Almost all fossil specimens are of spirally arranged clusters of the individuals' teeth, called "tooth whorls", which in life were embedded in the lower jaw. As with most extinct cartilaginous fish, the skeleton is mostly unknown. Fossils of Helicoprion are known from a 20 million year timespan during the Permian period from the Artinskian stage of the Cisuralian to the Roadian stage of the Guadalupian. The closest living relatives of Helicoprion are the chimaeras, though their relationship is very distant. The unusual tooth arrangement is thought to have been an adaption for feeding on soft bodied prey, and may have functioned as a deshelling mechanism for hard bodied cephalopods such as nautiloids and ammonoids. In 2013, systematic revision of Helicoprion via morphometric analysis of the tooth whorls found only H. davisii, H. bessonowi and H. ergassaminon to be valid, with some of the larger tooth whorls being outliers.
Erythrosuchidae are a family of large basal archosauriform carnivores that lived from the later Early Triassic (Olenekian) to the early Middle Triassic (Anisian).
Parareptilia ("near-reptiles") is a subclass or clade of basal sauropsids/reptiles, typically considered the sister taxon to Eureptilia. Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles, the first reptiles with advanced hearing systems, and the first large herbivorous reptiles. The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by the end of the period.
Tritylodontidae is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, with several mammalian traits including erect limbs, endothermy and details of the skeleton. They were the last-known family of the non-mammaliaform synapsids, persisting into the Early Cretaceous.
Garjainia is an extinct genus of erythrosuchid archosauriform reptile from the Olenekian of Russia and South Africa. It was approximately 1.5–2 metres (4.9–6.6 ft) long. It contained two species, Garjainia prima from the Yarengian/Yarkenskian Supergorizont of Russia, and Garjainia madiba from the Burgersdorp Formation of South Africa. "Vjuskovia triplicostata", a name assigned to some erythrosuchid fossils from Russia, has been synonymized with Garjainia prima.
Heterodontosauridae is a family of ornithischian dinosaurs that were likely among the most basal (primitive) members of the group. Their phylogenetic placement is uncertain but they are most commonly found to be primitive, outside of the group Genasauria. Although their fossils are relatively rare and their group small in numbers, they have been found on all continents except Australia and Antarctica, with a range spanning the Early Jurassic to the Early Cretaceous.
Omphalosaurus is an extinct genus of marine reptile from the Early Triassic to Middle Triassic, thought to be in the order of Ichthyosauria. Most of what is known about Omphalosaurus is based on multiple jaw fragments, ribs, and vertebrae. Specimens of Omphalosaurus have been described from the western United States, Poland, Austria and the island of Spitsbergen off the northern coast of Norway.
Brachyopoidea is a superfamily of temnospondyls that lived during the Mesozoic. It contains the families Brachyopidae and Chigutisauridae. The earliest records of brachyopids are from the Lower Triassic in Australia. The latest-surviving member of the superfamily is the chigutisaurid Koolasuchus from the Early Cretaceous of Australia.
The Eugeneodontida, sometimes also called Eugeneodontiformes, is an extinct and poorly known order of cartilaginous fishes. They possessed "tooth-whorls" on the symphysis of either the lower or both jaws and pectoral fins supported by long radials. They probably lacked pelvic fins and anal fins. The palatoquadrate was either fused to the skull or reduced. Now determined to be within the Holocephali, their closest living relatives are ratfish. The eugeneodonts are named after paleontologist Eugene S. Richardson, Jr. The Eugeneodontida disappeared in the Early Triassic. The geologically youngest fossils of the group are known from the Sulphur Mountain Formation, Vardebukta Formation and Wordie Creek Formation (Greenland).
Kuehneotherium is an early mammaliaform genus, previously considered a holothere, that lived during the Late Triassic-Early Jurassic Epochs and is characterized by reversed-triangle pattern of molar cusps. Although many fossils have been found, the fossils are limited to teeth, dental fragments, and mandible fragments. The genus includes Kuehneotherium praecursoris and all related species. It was first named and described by Doris M. Kermack, K. A. Kermack, and Frances Mussett in November 1967. The family Kuehneotheriidae and the genus Kuehneotherium were created to house the single species Kuehneotherium praecursoris. Modeling based upon a comparison of the Kuehneotherium jaw with other mammaliaforms indicates it was about the size of a modern-day shrew between 4 and 5.5 g at adulthood.
Clevosaurus is an extinct genus of rhynchocephalian reptile from the Late Triassic and the Early Jurassic periods. Species of Clevosaurus were widespread across Pangaea, and have been found on all continents except Australia and Antarctica. Five species of Clevosaurus have been found in ancient fissure fill deposits in south-west England and Wales, alongside other sphenodontians, early mammals and dinosaurs. In regards to its Pangaean distribution, C. hadroprodon is the oldest record of a sphenodontian from Gondwana, though its affinity to Clevosaurus has been questioned.
Microposaurus is an extinct genus of trematosaurid temnospondyl. Fossils are known from the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the Rouse Hill Siltstone of Australia that date back to the Anisian stage of the Middle Triassic. These aquatic creatures were the short snouted lineage from Trematosaurinae.
Helicoprionidae is an extinct, poorly known family of bizarre holocephalids within the poorly understood order Eugeneodontida. Members of the Helicoprionidae possessed a unique "tooth-whorl" on the symphysis of the lower jaw and pectoral fins supported by long radials. The closest living relatives of the Helicoprionidae and all other eugeneodontids are the ratfishes. The anatomy of the tooth-whorl differed amongst genus and species, some possessing complete spirals, others possessing halved spirals, and some with wedged half-spirals. Each tooth-whorl is thought to be adapted to a different type of prey, and a different predation strategy.
Wimanius is a genus of ichthyosaur from the Middle Triassic of Switzerland, containing a single species, Wimanius odontopalatus. It was described by Michael Maisch and Andreas Matzke in 1998 based on an incomplete skull from Monte San Giorgio, a mountain on the Swiss-Italian border. Wimanius possesses teeth on its palate, though whether they were located on the palatine or pterygoid is disputed. Other features of Wimanius include a large orbit and jugals with two rami of similar lengths. Different phylogenetic placements of Wimanius have been recovered by different studies, including it being a mixosaurid relative or a merriamosaur, and a monotypic family, Wimaniidae has been named for it. However, its validity has also been questioned, and synonymy with various other genera has been proposed.
Gephyrosaurus is a genus of early rhynchocephalian first described and named in 1980 by Susan E. Evans. They are distantly related to the extant Sphenodon with which they shared a number of skeletal features including a large tooth row along the side of the palatine bone and posterior process of the dentary bone. The type species, G. bridensis, lived during Early Jurassic in Wales, UK. Whiteside & Duffin (2017) described the second species, G. evansae, known from a partial maxilla recovered from Late Triassic (Rhaetian) fissure fills in Carboniferous Limestone in Somerset. Gephyrosaurus, other potential gephyrosaurids and Wirtembergia are the only rhynchocephalians to lie outside Sphenodontia in modern definitions of the group, and have been found to be more closely related to squamates in some phylogenetic analyses.
Fugusuchus is an extinct genus of archosauriform, probably the basal-most member of the family Erythrosuchidae. The genus is known from a single fossil from the middle Early Triassic Heshanggou Formation in Shanxi, China. The partial skeleton consists of an incomplete skull, parts of the right forelimb, and an intercentrum. The skeleton, known as GMB V 313, is currently in the Geological Museum of China in Beijing.
Hanosaurus is an extinct genus of marine reptiles that existed during the Triassic period in what is now China. The type species is Hanosaurus hupehensis. It was a small animal, with specimens measuring 79.4 cm (31.3 in) long in total body length, which likely fed on soft-bodied prey.
Wangisuchus is an extinct genus of archosauriform reptile from the Middle Triassic of China that is known from fragmentary fossil jaw bones. These bones were found at the Hsishihwa locality in the upper Ermaying Formation, which dates to the late Anisian stage about 242 million years ago. Wangisuchus was named in 1964 by Chinese paleontologist Yang Zhongjian, who described a single species, Wangisuchus tzeyii, on the basis of these bones.
