Caseodontidae Temporal range: Early Carboniferous to Early Triassic | |
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Life restoration of Caseodus , the type genus | |
The skull of Ornithoprion , an aberrant caseodont | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Chondrichthyes |
Subclass: | Holocephali |
Order: | † Eugeneodontida |
Clade: | † Caseodontoidea |
Family: | † Caseodontidae Zangerl, 1981 |
Type genus | |
Caseodus Zangerl, 1981 | |
Type species | |
Orodus basalis (=Caseodus basalis) Cope, 1894 | |
Genera | |
The Caseodontidae is an extinct family of eugeneodont holocephalans known from the late Paleozoic to earliest Mesozoic of Greenland, Canada and the United States. [1] [2] Members of the group are characterized by a reduced or absent palatoquadrate, elongate upper and mandibular rostra, and bulbous, crushing dentition, including a small symphyseal whorl of teeth on the lower jaw and batteries of teeth fused directly to the neurocranium. [3] [4] Several genera are known from partial or complete body fossils. [3] [5]
Unlike the distantly related helicoprionids, members of this family crossed the Permian-Triassic boundary and persisted into the Olenekian stage of the Early Triassic, [5] after which they became extinct. It is hypothesized that in life caseodonts fed on hard-shelled prey such as brachiopods due to their crushing tooth batteries, [4] and it has been proposed that the elongated rostra on the upper and lower jaws of some genera was an adaptation for prying prey off of the seabed. [6] Well preserved specimens are known from the Carboniferous of Nebraska and Indiana, [3] deposits in East Greenland, [2] and from the Sulphur Mountain Formation of British Columbia, which is the last known appearance of the group. [5]
Helicoprion is an extinct genus of shark-like eugeneodont fish. Almost all fossil specimens are of spirally arranged clusters of the individuals' teeth, called "tooth whorls", which in life were embedded in the lower jaw. As with most extinct cartilaginous fish, the skeleton is mostly unknown. Fossils of Helicoprion are known from a 20 million year timespan during the Permian period from the Artinskian stage of the Cisuralian to the Roadian stage of the Guadalupian. The closest living relatives of Helicoprion are the chimaeras, though their relationship is very distant. The unusual tooth arrangement is thought to have been an adaption for feeding on soft bodied prey, and may have functioned as a deshelling mechanism for hard bodied cephalopods such as nautiloids and ammonoids. In 2013, systematic revision of Helicoprion via morphometric analysis of the tooth whorls found only H. davisii, H. bessonowi and H. ergassaminon to be valid, with some of the larger tooth whorls being outliers.
Hybodus is an extinct genus of hybodont that lived from the Middle Triassic to the Late Cretaceous periods. Species closely related to the type species Hybodus reticulatus lived during the Early Jurassic epoch. Numerous species have been assigned to Hybodus spanning a large period of time, and it is currently considered a wastebasket taxon that is 'broadly polyphyletic' and requires reexamination.
Hesperosuchus is an extinct genus of crocodylomorph reptile that contains a single species, Hesperosuchus agilis. Remains of this pseudosuchian have been found in Late Triassic (Carnian) strata from Arizona and New Mexico. Because of similarities in skull and neck anatomy and the presence of hollow bones Hesperosuchus was formerly thought to be an ancestor of later carnosaurian dinosaurs, but based on more recent findings and research it is now known to be more closely related to crocodilians rather than dinosaurs.
Aphaneramma is an extinct genus of marine temnospondyl amphibian. It lived during the Early Triassic epoch, approximately 252–247 Ma ago, and had a global distribution.
Edestus is an extinct genus of eugeneodontid holocephalian fish known from the Late Carboniferous (Pennsylvanian) of the United Kingdom, Russia, and the United States. Most remains consist of isolated curved blades or "whorls" that are studded with teeth, that in life were situated within the jaws. Edestus is a Greek name derived from the word edeste, in reference to the aberrant quality and size of the species' teeth. The largest species, E. heinrichi, has been conservatively estimated to reach greater than 6.7 m (22 ft) in length, around the size of the largest known great white shark, possibly making it the largest marine predator to have ever existed up to that point.
Omphalosaurus is an extinct genus of marine reptile from the Early Triassic to Middle Triassic, thought to be in the order of Ichthyosauria. Most of what is known about Omphalosaurus is based on multiple jaw fragments, ribs, and vertebrae. Specimens of Omphalosaurus have been described from the western United States, Poland, Austria and the island of Spitsbergen off the northern coast of Norway.
Thalattosaurus meaning "sea lizard," from the Attic Greek thalatta (θάλαττα), "sea," and sauros (σαῦρος), "lizard," is an extinct genus of marine reptile in the family Thalattosauroidea. They were aquatic diapsids that are known exclusively from the Triassic period. It was a 2–3 metres (6.6–9.8 ft) long shellfish-eating reptile with paddle-like limbs and a down-turned rostrum occurring in the Lower and Middle Triassic Sulphur Mountain Formation of British Columbia as well as the Upper Triassic Hosselkus Limestone of California. It has gained notoriety as a result of studies on general diapsid phylogeny.
Fadenia is an extinct genus of eugeneodontid holocephalian chondrichthyan from the Carboniferous Period of Missouri, the Permian period of Greenland, and the Early Triassic epoch of British Columbia, Canada.
Caseodus is an extinct genus of eugeneodontid holocephalian from the Carboniferous of the United States and the Early Triassic of Canada. It was of medium size, measuring 1–1.5 m (3.3–4.9 ft) in length.
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like cartilaginous fish (chondrichthyans) which existed from the late Devonian to the Late Cretaceous. Hybodonts share a close common ancestry with modern sharks and rays (Neoselachii) as part of the clade Euselachii. They are distinguished from other chondrichthyans by their distinctive fin spines and cephalic spines present on the heads of males. An ecologically diverse group, they were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct.
The Eugeneodontida, sometimes also called Eugeneodontiformes, is an extinct and poorly known order of cartilaginous fishes. They possessed "tooth-whorls" on the symphysis of either the lower or both jaws and pectoral fins supported by long radials. They probably lacked pelvic fins and anal fins. The palatoquadrate was either fused to the skull or reduced. Now determined to be within the Holocephali, their closest living relatives are chimaeras. The eugeneodonts are named after paleontologist Eugene S. Richardson, Jr. The group first appeared in the fossil record during the late Mississippian (Serpukhovian). The youngest eugeneodonts are known from the Early Triassic. The geologically youngest fossils of the group are known from the Sulphur Mountain Formation, Vardebukta Formation and Wordie Creek Formation (Greenland).
Australosomus is an extinct genus of prehistoric ray-finned fish that lived during the Early Triassic epoch in what is now Greenland, Kenya, Tanzania, Madagascar, South Africa and Canada.
Helmolepis is an extinct genus of ray-finned fish that lived during the Early Triassic epoch in what is now Greenland, Madagascar and Canada. Species of Helmolepis are small. This genus is closely related with Platysiagum.
Anaschisma is an extinct genus of large temnospondyls. These animals were part of the family called Metoposauridae, which filled the crocodile-like predatory niches in the late Triassic. It had a large skull about 62 centimetres (24 in) long, and possibly reached 3 metres (9.8 ft) long. It was an ambush hunter, snapping up anything small enough to fit in its huge jaws. It was very common during the Late Triassic in what is now the American Southwest.
Bobasatraniiformes is an extinct order of durophagous ray-finned fish that existed from the late Permian to the Middle Triassic in both marine and freshwater environments. The order includes two families: Bobasatraniidae, with the genera Bobasatrania, Ebenaqua, and Ecrinesomus, and Dorypteridae, comprising only the genus Dorypterus (monotypy). Bobasatraniiformes had a somewhat global distribution; fossils are found in Africa (Madagascar), Asia (Pakistan), Australia, Europe, and North America.
Ymeria is an extinct genus of early stem tetrapod from the Devonian of Greenland. Of the two other genera of stem tetrapods from Greenland, Acanthostega and Ichthyostega, Ymeria is most closely related to Ichthyostega, though the single known specimen is smaller, the skull about 10 cm in length. A single interclavicle resembles that of Ichthyostega, an indication Ymeria may have resembled this genus in the post-cranial skeleton.
Ornithoprion is a monotypic genus of extinct eugeneodont holocephalan in the family Caseodontidae. The type and only species, O. hertwigi, lived during the Moscovian stage of the Carboniferous, between 315.2 to 307 million years ago, and is known from black shale deposits in what is now the Midwestern United States. The discovery and description of Ornithoprion, performed primarily via radiography, helped clarify the cranial anatomy of the eugeneodonts, a group which were previously known primarily from isolated tooth whorls. The genus derives its name from the elongate, bill-like rostrum and large eyes, which vaguely resemble the features of a bird, while the species name honors Oscar Hertwig. It is known from specimens preserving the skull and anterior portion of the body.
Barbclabornia is an extinct genus of xenacanth from the Early Permian and possibly upper Pennsylvanian of North America. The genus contains a single described species: B. luedersensis. It has been found in several places within Asselian and Atinskian formations, including the Clear Fork, Albany, Wichita, and Dunkard Groups. There are possible examples from the Gzhelian-aged Admire, Monongahela, and Conemaugh groups.
The Eugeneodontidae is an extinct family of eugeneodont holocephalans known from the late Paleozoic the United States and Iran. A defining trait of the group is pavement-like dentition with blade-like teeth, implying a divergent diet to the closely related caseodontids which they otherwise closely resemble. Both Eugeneodus and Gilliodus are known from virtually complete skeletons, including extensive patches of preserved dermal denticles. The name of both the family and the type genus, and by extension the entire order of Eugeneodontida, are in honor of Paleontologist Eugene S. Richardson Jr.
Paredestus is a monotypic genus of extinct eugeneodont holocephalan from the Early Triassic of Canada. The type and only species, P. bricircum represents the last known member of the superfamily Edestoidea and among the last known eugeneodonts. It was named in 2008 based on tooth and jaw material, with the holotype representing the only known specimen.
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