Listracanthus | |
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Denticles of the holotype specimen of L. pectenatus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Chondrichthyes |
Order: | incertae sedis |
Family: | † Listracanthidae |
Genus: | † Listracanthus Newberry & Worthen, 1870 |
Species [1] | |
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Listracanthus is a genus of extinct chondrichthyan with uncertain affinities. Species of Listracanthus are known primarily from their tremendous, feather-like denticles, which range up to four inches in length. The denticles had a large main spine, from which secondary spines emanate from the sides, like the barbs of a feather or a comb. Listracanthus first appeared in late Carboniferous strata in North America, and eventually disappear from the fossil record some time during the Early Triassic. [1]
The appearance of these sharks are largely unknown. However, author and illustrator Ray Troll mentions in his book, Sharkabet, about how paleontologist Rainer Zangerl once discovered a large shale slab containing a long, eel-like fish covered in long, spine-like denticles characteristic of the genus, only to have it dry out and crumble into dust. As such, according to Zangerl's account, Troll reconstructs Listracanthus as resembling a tremendous, fiercely bristled frill shark. [3]
Martill et al., (2014) created the genus Acanthorhachis for the species formerly known as "Listracanthus" spinatus (Bolton, 1896). They also erected the family Listracanthidae to encompass the two genera. [4]
Chondrichthyes is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or bony fish, which have skeletons primarily composed of bone tissue. Chondrichthyes are aquatic vertebrates with paired fins, paired nares, placoid scales, conus arteriosus in the heart, and a lack of opercula and swim bladders. Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates.
Chimaeras are cartilaginous fish in the order Chimaeriformes, known informally as ghost sharks, rat fish, spookfish, or rabbit fish; the last three names are not to be confused with rattails, Opisthoproctidae, or Siganidae, respectively.
Cladoselache is an extinct genus of shark-like chondrichthyan from the Late Devonian (Famennian) of North America. It was similar in body shape to modern lamnid sharks, but was not closely related to lamnids or to any other modern (selachian) shark. As an early chondrichthyan, it had yet to evolve traits of modern sharks such as accelerated tooth replacement, a loose jaw suspension, enameloid teeth, and possibly claspers.
Helicoprion is an extinct genus of shark-like eugeneodont fish. Almost all fossil specimens are of spirally arranged clusters of the individuals' teeth, called "tooth whorls", which in life were embedded in the lower jaw. As with most extinct cartilaginous fish, the skeleton is mostly unknown. Fossils of Helicoprion are known from a 20 million year timespan during the Permian period from the Artinskian stage of the Cisuralian to the Roadian stage of the Guadalupian. The closest living relatives of Helicoprion are the chimaeras, though their relationship is very distant. The unusual tooth arrangement is thought to have been an adaption for feeding on soft bodied prey, and may have functioned as a deshelling mechanism for hard bodied cephalopods such as nautiloids and ammonoids. In 2013, systematic revision of Helicoprion via morphometric analysis of the tooth whorls found only H. davisii, H. bessonowi and H. ergassaminon to be valid, with some of the larger tooth whorls being outliers.
Stethacanthus is an extinct genus of shark-like cartilaginous fish which lived from the Late Devonian to Late Carboniferous epoch, dying out around 298.9 million years ago. Fossils have been found in Australia, Asia, Europe and North America.
Xenacanthida is an order or superorder of extinct shark-like chondrichthyans known from the Carboniferous to Triassic. They were native to freshwater, marginal marine and shallow marine habitats. Some xenacanths may have grown to lengths of 5 m (16 ft). Most xenacanths died out at the end of the Permian in the End-Permian Mass Extinction, with only a few forms surviving into the Triassic.
Hybodus is an extinct genus of hybodont that lived from the Middle Triassic to the Late Cretaceous periods. Species closely related to the type species Hybodus reticulatus lived during the Early Jurassic epoch. Numerous species have been assigned to Hybodus spanning a large period of time, and it is currently considered a wastebasket taxon that is 'broadly polyphyletic' and requires reexamination.
Tristychius is an extinct genus of euselachian chondrichthyan from the Carboniferous period (Visean). Fossils of T. arcuatus, the type and only species, including fin spines have been found in Scotland.
Edestus is an extinct genus of eugeneodontid holocephalian fish known from the Late Carboniferous (Pennsylvanian) of the United Kingdom, Russia, and the United States. Most remains consist of isolated curved blades or "whorls" that are studded with teeth, that in life were situated within the midline of the upper and lower jaws. Edestus is a Greek name derived from the word edeste, in reference to the aberrant quality and size of the species' teeth. The largest species, E. heinrichi, has been conservatively estimated to reach greater than 6.7 m (22 ft) in length, around the size of the largest known great white shark, possibly making it the largest marine predator to have ever existed up to that point.
Falcatus is an extinct genus of falcatid chondrichthyan which lived during the early Carboniferous Period in Bear Gulch bay in what is now Montana.
Stethacanthidae is an extinct family of prehistoric holocephalians. It is estimated to have existed approximately between 380 and 300 million years ago. Members of this family are noted for their peculiar dorsal fin.
Iniopteryx is an extinct genus of cartilaginous fish. It is from the Pennsylvanian epoch of the Carboniferous period, approximately 300 million years ago. Their fossils have been found in North America, primarily in two states: Ohio and Montana. In general, very little is known about this species.
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like cartilaginous fish (chondrichthyans) which existed from the late Devonian to the Late Cretaceous. Hybodonts share a close common ancestry with modern sharks and rays (Neoselachii) as part of the clade Euselachii. They are distinguished from other chondrichthyans by their distinctive fin spines and cephalic spines present on the heads of males. An ecologically diverse group, they were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct.
Squaloraja is an extinct genus of ray-like marine chimaeriform fish from the Early Jurassic of Europe.
Ornithoprion is a monotypic genus of extinct eugeneodont holocephalan in the family Caseodontidae. The type and only species, O. hertwigi, lived during the Moscovian stage of the Carboniferous, between 315.2 to 307 million years ago, and is known from black shale deposits in what is now the Midwestern United States. The discovery and description of Ornithoprion, performed primarily via radiography, helped clarify the anatomy of the eugeneodonts, a group which were previously known primarily from isolated tooth whorls. The genus derives its name from the elongate, bill-like rostrum and large eyes, which vaguely resemble the features of a bird, while the species name honors Oscar Hertwig. It is known from specimens preserving the skull and anterior portion of the body.
Antarctilamna is an extinct genus of Devonian cartilaginous fish originally exemplified by Antarctilamna prisca from South Eastern Australia and Antarctica. The latest occurring described species is Antarctilamna ultima from the Waterloo Farm lagerstätte in South Africa. Antarctilamna has robust ctenacanthid-like spines which lack a deep insertion area, and are borne in front of the first dorsal fin; in addition to distinctive diplodont teeth with small intermediate cusps. Antarctilamna-like spines, known from the Bunga Beds locality in Australia have been ascribed to A. prisca.
Acanthorhachis is an enigmatic extinct genus of chondrichthyan from the Carboniferous period. Its name is derived from the Greek word acanthos meaning "spine" and the Greek suffix for spine, -rhachis. This is due to the spine-like dermal denticles and their subsidiary spines, which coated the exterior of the animal. The authors who erected this genus suggested the common name "The Spiny Spined Shark." The type species Listracanthus spinatus was in 1896 named by Herbert Bolton.
Arctacanthus is an extinct genus of cartilaginous fish from the Permian and possibly Triassic periods.
Bandringa is an extinct genus of elasmobranch known from the Pennsylvanian subperiod of the Carboniferous period that was part of the monotypic family Bandringidae. There is currently a single known species, B. rayi, described in 1969. It is known from exceptionally preserved individuals found in the Mazon Creek Lagerstätte of Illinois which dates back to the late Moscovian stage.
Paredestus is a monotypic genus of extinct eugeneodont holocephalan from the Early Triassic of Canada. The type and only species, P. bricircum represents the last known member of the superfamily Edestoidea and among the last known eugeneodonts. It was named in 2008 based on tooth and jaw material, with the holotype representing the only known specimen.