Lake zones |
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Lake stratification |
Lake types |
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Lake stratification is the tendency of lakes to form separate and distinct thermal layers during warm weather. Typically stratified lakes show three distinct layers: the epilimnion, comprising the top warm layer; the thermocline (or metalimnion), the middle layer, whose depth may change throughout the day; and the colder hypolimnion, extending to the floor of the lake.
Every lake has a set mixing regime that is influenced by lake morphometry and environmental conditions. However, changes to human influences in the form of land use change, increases in temperature, and changes to weather patterns have been shown to alter the timing and intensity of stratification in lakes around the globe. [1] [2] Rising air temperatures have the same effect on lake bodies as a physical shift in geographic location, with tropical zones being particularly sensitive. [2] [1] These changes can further alter the fish, zooplankton, and phytoplankton community composition, in addition to creating gradients that alter the availability of dissolved oxygen and nutrients. [3] [4]
The thermal stratification of lakes refers to a change in the temperature at different depths in the lake, and is due to the density of water varying with temperature. [5] Cold water is denser than warm water and the epilimnion generally consists of water that is not as dense as the water in the hypolimnion. [6] However, the temperature of maximum density for freshwater is 4 °C. In temperate regions where lake water warms up and cools through the seasons, a cyclical pattern of overturn occurs that is repeated from year to year as the cold dense water at the top of the lake sinks (see stable and unstable stratification). For example, in dimictic lakes the lake water turns over during the spring and the fall. This process occurs more slowly in deeper water and as a result, a thermal bar may form. [5] If the stratification of water lasts for extended periods, the lake is meromictic.
Heat is transported very slowly between the mixed layers of a stratified lake, where the diffusion of heat just one vertical meter takes about a month. The interaction between the atmosphere and lakes depends on how solar radiation is distributed, which is why water turbulence, mainly caused by wind stress, can greatly increase the efficiency of heat transfer. [7] In shallow lakes, stratification into epilimnion, metalimnion, and hypolimnion often does not occur, as wind or cooling causes regular mixing throughout the year. These lakes are called polymictic. There is not a fixed depth that separates polymictic and stratifying lakes, as apart from depth, this is also influenced by turbidity, lake surface area, and climate. [8]
The lake mixing regime (e.g. polymictic, dimictic, meromictic) [9] describes the yearly patterns of lake stratification that occur in most years. However, short-term events can influence lake stratification as well. Heat waves can cause periods of stratification in otherwise mixed, shallow lakes, [10] while mixing events, such as storms or large river discharge, can break down stratification. [11] Weather conditions induce a more rapid response in larger, shallower lakes, so these lakes are more dynamic and less understood. However, mixing regimes that are known to exist in large, shallow lakes are mostly diurnal, and the stratification is easily disturbed. Lake Taihu in China is an example of a large, shallow, diurnal lake, where even though the depth does not reach more than 3 metres (9.8 ft), the lake’s water turbidity is still dynamic enough to stratify and de-stratify due to the absorption of solar radiation mostly in the upper layer. [12] The tendency for stratification to become disrupted affects the rate of transport and consumption of nutrients, in turn affecting the presence of algal growth. [13] Stratification and mixing regimes in Earth’s largest lakes are also poorly understood, yet changes in thermal distributions, such as the rising temperatures found over time in Lake Michigan’s deep waters, have the ability to significantly alter the largest freshwater ecosystems on the planet. [14]
Recent research suggests that seasonally ice-covered dimictic lakes may be described as "cryostratified" or "cryomictic" according to their wintertime stratification regimes. [15] Cryostratified lakes exhibit inverse stratification near the ice surface and have depth-averaged temperatures near 4°C, while cryomictic lakes have no under-ice thermocline and have depth-averaged winter temperatures closer to 0°C. [16]
Circulation processes during mixing periods cause the movement of oxygen and other dissolved nutrients, distributing them throughout the body of water. [7] In lakes where benthic organisms are prominent, the respiration and consumption of these bottom-feeders may outweigh the mixing properties of strongly stratified lakes, resulting in zones of extremely low near-bottom oxygen and nutrient concentrations. This can be harmful to benthic organisms such as shellfish, which in the worst cases can wipe out entire populations. [17] The accumulation of dissolved carbon dioxide in three meromictic lakes in Africa (Lake Nyos and Lake Monoun in Cameroon and Lake Kivu in Rwanda) is potentially dangerous because if one of these lakes is triggered into limnic eruption, a very large quantity of carbon dioxide can quickly leave the lake and displace the oxygen needed for life by people and animals in the surrounding area.
In temperate latitudes, many lakes that become stratified during the summer months de-stratify during cooler windier weather with surface mixing by wind being a significant driver in this process. This is often referred to as "autumn turn-over". The mixing of the hypolimnium into the mixed water body of the lake recirculates nutrients, particularly phosphorus compounds, trapped in the hypolimnion during the warm weather. It also poses a risk of oxygen sag as a long established hypolimnion can be anoxic or very low in oxygen.
Lake mixing regimes can shift in response to increasing air temperatures. Some dimictic lakes can turn into monomictic lakes, while some monomictic lakes might become meromictic, as a consequence of rising temperatures. [18]
Many types of aeration equipment have been used to thermally de-stratify lakes, particularly lakes subject to low oxygen or undesirable algal blooms. [19] In fact, natural resource and environmental managers are often challenged by problems caused by lake and pond thermal stratification. [6] [20] [21] Fish die-offs have been directly associated with thermal gradients, stagnation, and ice cover. [22] Excessive growth of plankton may limit the recreational use of lakes and the commercial use of lake water. With severe thermal stratification in a lake, the quality of drinking water also can be adversely affected. [6] For fisheries managers, the spatial distribution of fish within a lake is often adversely affected by thermal stratification and in some cases may indirectly cause large die-offs of recreationally important fish. [22] One commonly used tool to reduce the severity of these lake management problems is to eliminate or lessen thermal stratification through water aeration. [20] Aeration has met with some success, although it has rarely proved to be a panacea. [21]
Every lake has a set mixing regime that is influenced by lake morphometry and environmental conditions. However, changes to human influences in the form of land use change, increases in temperatures, and changes to weather patterns have been shown to alter the timing and intensity of stratification in lakes around the globe. [1] [2] These changes can further alter the fish, zooplankton, and phytoplankton community composition, in addition to creating gradients that alter the availability of dissolved oxygen and nutrients. [3] [4]
There are a number of ways in which changes in human land use influence lake stratification and consequently water conditions. Urban expansion has led to the construction of roads and houses close to previously isolated lakes, sometimes causing increased runoff and pollution. The addition of particulate matter to lake bodies can lower water clarity, resulting in stronger thermal stratification and overall lower average water column temperatures, which can eventually affect the onset of ice cover. [23] Water quality can also be influenced by the runoff of salt from roads and sidewalks, which often creates a benthic saline layer that interferes with vertical mixing of surface waters. [4] Further, the saline layer can prevent dissolved oxygen from reaching the bottom sediments, decreasing phosphorus recycling and affecting microbial communities. [4]
On a global scale, rising temperatures and changing weather patterns can also affect stratification in lakes. Rising air temperatures have the same effect on lake bodies as a physical shift in geographic location, with tropical zones being particularly sensitive. [2] [1] The intensity and scope of impact depends on location and lake morphometry, but in some cases can be so extreme as to require a reclassification from monomictic to dimictic (e.g. Great Bear Lake). [2] Globally, lake stratification appears to be more stable with deeper and steeper thermoclines, and average lake temperature as a main determinant in the stratification response to changing temperatures. [1] Further, surface warming rates are much greater than bottom warming rates, again indicating stronger thermal stratification across lakes. [1]
Changes to stratification patterns can also alter the community composition of lake ecosystems. In shallow lakes, temperature increases can alter the diatom community; while in deep lakes, the change is reflected in the deep chlorophyll layer taxa. [3] Changes in mixing patterns and increased nutrient availability can also affect zooplankton species composition and abundance, while decreased nutrient availability can be detrimental for benthic communities and fish habitat. [3] [4]
In northern temperate lakes, as climate change continues to cause increased variability in weather patterns as well as the timing of ice-on and ice-off dates, subsequent changes in stratification patterns from year to year can also have impacts across multiple trophic levels. [24] [25] [26] Fluctuations in stratification consistency can accelerate deoxygenation of lakes, nutrient mineralization, and phosphorus release, having significant consequences for phytoplankton species. [26] [27] Furthermore, these changes in phytoplankton species composition and abundance can lead to adverse effects on fish recruitment, such as walleye. When these asynchronies in predator and prey populations occur year after year due to changes in stratification, populations may take years to rebound to their “normal” consistency. [27] Combined with typically warmer lake temperatures associated with stratification patterns brought on by climate change, variable prey populations from year-to-year can be detrimental to cold water fish species. [28]
Limnology is the study of inland aquatic ecosystems. The study of limnology includes aspects of the biological, chemical, physical, and geological characteristics of fresh and saline, natural and man-made bodies of water. This includes the study of lakes, reservoirs, ponds, rivers, springs, streams, wetlands, and groundwater. Water systems are often categorized as either running (lotic) or standing (lentic).
The hypolimnion or under lake is the dense, bottom layer of water in a thermally-stratified lake. The word "hypolimnion" is derived from Ancient Greek: λιμνίον, romanized: limníon, lit. 'lake'. It is the layer that lies below the thermocline.
The epilimnion or surface layer is the top-most layer in a thermally stratified lake.
A thermocline is a distinct layer based on temperature within a large body of fluid with a high gradient of distinct temperature differences associated with depth. In the ocean, the thermocline divides the upper mixed layer from the calm deep water below.
A meromictic lake is a lake which has layers of water that do not intermix. In ordinary, holomictic lakes, at least once each year, there is a physical mixing of the surface and the deep waters.
The spring bloom is a strong increase in phytoplankton abundance that typically occurs in the early spring and lasts until late spring or early summer. This seasonal event is characteristic of temperate North Atlantic, sub-polar, and coastal waters. Phytoplankton blooms occur when growth exceeds losses, however there is no universally accepted definition of the magnitude of change or the threshold of abundance that constitutes a bloom. The magnitude, spatial extent and duration of a bloom depends on a variety of abiotic and biotic factors. Abiotic factors include light availability, nutrients, temperature, and physical processes that influence light availability, and biotic factors include grazing, viral lysis, and phytoplankton physiology. The factors that lead to bloom initiation are still actively debated.
Thermal pollution, sometimes called "thermal enrichment", is the degradation of water quality by any process that changes ambient water temperature. Thermal pollution is the rise or drop in the temperature of a natural body of water caused by human influence. Thermal pollution, unlike chemical pollution, results in a change in the physical properties of water. A common cause of thermal pollution is the use of water as a coolant by power plants and industrial manufacturers. Urban runoff—stormwater discharged to surface waters from rooftops, roads, and parking lots—and reservoirs can also be a source of thermal pollution. Thermal pollution can also be caused by the release of very cold water from the base of reservoirs into warmer rivers.
A lake ecosystem or lacustrine ecosystem includes biotic (living) plants, animals and micro-organisms, as well as abiotic (non-living) physical and chemical interactions. Lake ecosystems are a prime example of lentic ecosystems, which include ponds, lakes and wetlands, and much of this article applies to lentic ecosystems in general. Lentic ecosystems can be compared with lotic ecosystems, which involve flowing terrestrial waters such as rivers and streams. Together, these two ecosystems are examples of freshwater ecosystems.
Ocean stratification is the natural separation of an ocean's water into horizontal layers by density, which is generally stable because warm water floats on top of cold water, and heating is mostly from the sun, which reinforces that arrangement. Stratification is reduced by wind-forced mechanical mixing, but reinforced by convection. Stratification occurs in all ocean basins and also in other water bodies. Stratified layers are a barrier to the mixing of water, which impacts the exchange of heat, carbon, oxygen and other nutrients. The surface mixed layer is the uppermost layer in the ocean and is well mixed by mechanical (wind) and thermal (convection) effects. Climate change is causing the upper ocean stratification to increase.
A dimictic lake is a body of freshwater whose difference in temperature between surface and bottom layers becomes negligible twice per year, allowing all strata of the lake's water to circulate vertically. All dimictic lakes are also considered holomictic, a category which includes all lakes which mix one or more times per year. During winter, dimictic lakes are covered by a layer of ice, creating a cold layer at the surface, a slightly warmer layer beneath the ice, and a still-warmer unfrozen bottom layer, while during summer, the same temperature-derived density differences separate the warm surface waters, from the colder bottom waters. In the spring and fall, these temperature differences briefly disappear, and the body of water overturns and circulates from top to bottom. Such lakes are common in mid-latitude regions with temperate climates.
Monomictic lakes are holomictic lakes that mix from top to bottom during one mixing period each year. Monomictic lakes may be subdivided into cold and warm types.
Amictic lakes are "perennially sealed off by ice, from most of the annual seasonal variations in temperature." Amictic lakes exhibit inverse cold water stratification whereby water temperature increases with depth below the ice surface 0 °C (less-dense) up to a theoretical maximum of 4 °C.
Polymictic lakes are holomictic lakes that are too shallow to develop thermal stratification; thus, their waters can mix from top to bottom throughout the ice-free period. Polymictic lakes can be divided into cold polymictic lakes, and warm polymictic lakes. While such lakes are well-mixed on average, during low-wind periods, weak and ephemeral stratification can often develop.
The Trophic State Index (TSI) is a classification system designed to rate water bodies based on the amount of biological productivity they sustain. Although the term "trophic index" is commonly applied to lakes, any surface water body may be indexed.
A lake is an often naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from the ocean, although they may be connected with the ocean by rivers, such as Lake Ontario. Most lakes are freshwater and account for almost all the world's surface freshwater, but some are salt lakes with salinities even higher than that of seawater. Lakes vary significantly in surface area and volume of water.
Euxinia or euxinic conditions occur when water is both anoxic and sulfidic. This means that there is no oxygen (O2) and a raised level of free hydrogen sulfide (H2S). Euxinic bodies of water are frequently strongly stratified; have an oxic, highly productive, thin surface layer; and have anoxic, sulfidic bottom water. The word "euxinia" is derived from the Greek name for the Black Sea (Εὔξεινος Πόντος (Euxeinos Pontos)) which translates to "hospitable sea". Euxinic deep water is a key component of the Canfield ocean, a model of oceans during part of the Proterozoic eon (a part specifically known as the Boring Billion) proposed by Donald Canfield, an American geologist, in 1998. There is still debate within the scientific community on both the duration and frequency of euxinic conditions in the ancient oceans. Euxinia is relatively rare in modern bodies of water, but does still happen in places like the Black Sea and certain fjords.
Lake metabolism represents a lake's balance between carbon fixation and biological carbon oxidation. Whole-lake metabolism includes the carbon fixation and oxidation from all organism within the lake, from bacteria to fishes, and is typically estimated by measuring changes in dissolved oxygen or carbon dioxide throughout the day.
Stratification in water is the formation in a body of water of relatively distinct and stable layers by density. It occurs in all water bodies where there is stable density variation with depth. Stratification is a barrier to the vertical mixing of water, which affects the exchange of heat, carbon, oxygen and nutrients. Wind-driven upwelling and downwelling of open water can induce mixing of different layers through the stratification, and force the rise of denser cold, nutrient-rich, or saline water and the sinking of lighter warm or fresher water, respectively. Layers are based on water density: denser water remains below less dense water in stable stratification in the absence of forced mixing.
Lake Lacawac is located at the very middle of Lacawac's Sanctuary Field Station in Pennsylvania and has been deemed the "southernmost unpolluted glacial lake in North America." Lake Lacawac has proven to be invaluable to researchers and students to conduct field experiments in order to learn more about the limnology of the lake.
An alpine lake is a high-altitude lake in a mountainous area, usually near or above the tree line, with extended periods of ice cover. These lakes are commonly glacial lakes formed from glacial activity but can also be formed from geological processes such as volcanic activity or landslides. Many alpine lakes that are fed from glacial meltwater have the characteristic bright turquoise green color as a result of glacial flour, suspended minerals derived from a glacier scouring the bedrock. When active glaciers are not supplying water to the lake, such as a majority of Rocky Mountains alpine lakes in the United States, the lakes may still be bright blue due to the lack of algal growth resulting from cold temperatures, lack of nutrient run-off from surrounding land, and lack of sediment input. The coloration and mountain locations of alpine lakes attract lots of recreational activity.