Stipatosteus Temporal range: Lower Devonian | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | † Placodermi |
Order: | † Arthrodira |
Family: | † Phlyctaeniidae |
Genus: | † Stipatosteus Plax & Newman, 2020 |
Species | |
†S. svidunovitchiPlax & Newman, 2020 |
Stipatosteus is an extinct genus of phlyctaeniid arthrodire from the Lower Devonian of Belarus. The genus is only known from one species, being S. svidunovitchi from the Lepel Beds. [1] Stipatosteus was the first Phlyctaenioideid to be found in the Devonian of Berlarus with only one other, Valentinaspis , being described since. [2]
The material that would be later described as Stipatosteus is from the Korma 1 borehole drilled near the Zhlobin Saddle at depths of between 340.8 and 337.7 meters. These remains were collected with the remains of multiple species Acanthodians along with many pieces from indeterminate vertebrates and small fragments of invertebrates. [1]
Stipatosteus is estimated to only be around 40-50 mm and is only known from four pieces of the trunk armor along with a few less complete fragments. Most of these plates have tubercles with star-shaped bases which are arranged in tightly-packed rows. These tubercles would have grown on top of one another as the animal aged causing some seen on the armor to have curving and splitting ridges moving outwards from the center. The spinal plates on the sides of the trunk are long and covered in rounded tubercles with only half of the spine extending out from where they attach to the anterior ventrolateral plate. There seems to be a lack of ornamentation on the anterior ventrolateral plate from the described remains, being the only part where in doesn't appear. These plates are very short when compared to other such as Phlyctaenius , being shorter than the posterior ventrolateral plates. [1]
The trunk armor of Stipatosteus is similar to a few different groups of placoderms such as that seen in Actinolepinans though lacks anterior ventral plate. This trait, along with the length of the spinal plates, put the genus within the family Phlyctaeniidae. [1]
Placoderms are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates, and the rest of the body was scaled or naked depending on the species.
Mizia is a genus of primitive antiarch placoderm found in Emsian-aged marine strata of Early Devonian China.
Dunkleosteus is an extinct genus of large arthrodire ("jointed-neck") fish that existed during the Late Devonian period, about 382–358 million years ago. It was a pelagic fish inhabiting open waters, and one of the first vertebrate apex predators of any ecosystem.
Bothriolepis was a widespread, abundant and diverse genus of antiarch placoderms that lived during the Middle to Late Devonian period of the Paleozoic Era. Historically, Bothriolepis resided in an array of paleo-environments spread across every paleocontinent, including near shore marine and freshwater settings. Most species of Bothriolepis were characterized as relatively small, benthic, freshwater detritivores, averaging around 30 centimetres (12 in) in length. However, the largest species, B. rex, had an estimated bodylength of 170 centimetres (67 in). Although expansive with over 60 species found worldwide, comparatively Bothriolepis is not unusually more diverse than most modern bottom dwelling species around today.
Groenlandaspis is an extinct genus of arthrodire from the Late Devonian. Fossils of the different species are found in late Devonian strata in all continents except eastern Asia. The generic name commemorates the fact that the first specimens of the type species were found in Greenland.
Cheirolepis is an extinct genus of marine and freshwater ray-finned fish that lived in the Devonian period of Europe and North America. It is the only genus yet known within the family Cheirolepididae and the order Cheirolepidiformes. It was among the most basal of the Devonian actinopterygians and is considered the first to possess the "standard" dermal cranial bones seen in later actinopterygians.
Rhenanida is an order of scaly placoderms. Unlike most other placoderms, the rhenanids' armor was made up of a mosaic of unfused scales and tubercles. The patterns and components of this "mosaic" correspond to the plates of armor in other, more advanced placoderms, suggesting that the ancestral placoderm had armor made of unfused components, as well.
Lunaspis is an extinct genus of armor-plated petalichthyid placoderm fish that lived in shallow marine environments of the Early Devonian period, from approximately 409.1 to 402.5 million year ago. Fossils have been found in Germany, China and Australia. There are three different identified species of within the genus Lunaspis: L. broilii, L. heroldi, and L. prumiensis.
Phyllolepis is the type genus of Phyllolepida, an extinct taxon of arthrodire placoderm fish from the middle to late Devonian. The species of Phyllolepis, themselves, are restricted to the Famennian-aged freshwater strata of the Late Devonian, around 360 million years ago. Fossils of this genus have been found primarily in Europe and North America. The end of the Devonian saw them disappear in a mass extinction.
Gaspeaspis is an extinct genus of placoderm fish, which lived during the Middle Devonian period of North America.
Holonema is an extinct genus of relatively large, barrel-shaped arthrodire placoderms that were found in oceans throughout the world from the Mid to Late Devonian, when the last species perished in the Frasnian-Fammian extinction event. Most species of the genus are known from fragments of their armor, but the Gogo Reef species, H. westolli, is known from whole, articulated specimens.
Holonematidae is an extinct family of relatively large arthrodire placoderms from the Early to Late Devonian. Almost all fossil specimens are of armor fragments, though, all have distinctive ornamentation, often of unique arrangements and patterns of tubercles, that are diagnostic of the family. The trunkshield is very elongated, giving the armor an overall "barrel" like appearance.
Buchanosteidae is a family of arthrodire placoderms that lived from the Early to Middle Devonian. Fossils appear in various strata in Russia, Central Asia, Australia, and China.
Microbrachiidae is an extinct family of tiny, advanced antiarch placoderms closely related to the bothriolepidids.
Oxyosteus is a genus of trout-sized, highly compressed arthrodire placoderms from the Late Devonian of Europe: The two described species are restricted to the Late Frasnian-aged Kellwasserkalk Fauna of Bad Wildungen, while a median dorsal plate of an unnamed species is known from the Middle Frasnian Holy Cross Mountains of Poland.
Microbrachius is an extinct genus of tiny, advanced antiarch placoderms closely related to the bothriolepids. Specimens range in age from the Lower Devonian Late Emsian Stage to the Middle Devonian Upper Givetian Stage. They are characterized by having large heads with short thoracic armor of an average length of 2–4 cm. There are patterns of small, but noticeable tubercles on the armor, with the arrangement varying from species to species. Specimens of Microbrachius have been found in Scotland, Belarus, Estonia, and China.
Asterolepis is an extinct genus of antiarch placoderms from the Devonian of North and South America and Europe. They were heavily armored flat-headed benthic detritivores with distinctive jointed limb-like pectoral fins and hollow spine. The armor plate gives the Asterolepis a box-like shape. Its pectoral fins are also armored but the caudal and dorsal fin are not. The first fossils were named by M. Eichwald in 1840 after noticing star-like markings on the fossils.
Qilinyu is a genus of early placoderm from the late Silurian of China. It contains a single species, Qilinyu rostrata, from the Xiaoxiang fauna of the Kuanti Formation. Along with its contemporary Entelognathus, Qilinyu is an unusual placoderm showing some traits more similar to bony fish, such as dermal jaw bones and lobe-like fins. It can be characterized by adaptations for a benthic lifestyle, with the mouth and nostrils on the underside of the head, similar to the unrelated antiarch placoderms. The shape of the skull has been described as "dolphin-like", with a domed cranium and a short projecting rostrum.
Actinolepis is an extinct genus of actinolepid placoderm from the Early Devonian. Four species are known: A. magna from Estonia, A. spinosa from Latvia, the type species A. tuberculata from New Zealand and A. zaikai from Belarus.
Kimbryanodus is a genus of extinct ptyctodontid placoderm fish from the Frasnian of Australia.These placoderms can be told apart from others due to the large eyes, crushing tooth plates, long bodies, reduced armor, and a superficial resemblance to holocephalid fish. The group is so far the only Placoderms known with sexually dimorphic features. The fossils occur as small three dimensional isolated plates. Because of these new specimens the Ptyctodontid grouping got a taxonomic classification, it found that the genus Rhamphodopsis to be the most basal taxa. They are divided by having the more basal taxa having a median dorsal spine, a simple spinal plate, and a simple V-shaped overlap of the anterior lateral and the anterior dorsolateral plates.