Ursoidea

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Ursoidea
Temporal range: Bartonian - Present
37–0  Ma
Cephalogale shareri.jpg
Life reconstruction of Cephalogale shareri
Brown bear (Ursus arctos), Viiksimo, Kainuu region, Finland, 16 June 2018 (43094873292).jpg
Brown bear (Ursus arctos)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Parvorder: Ursida
Superfamily: Ursoidea
Fischer von Waldheim, 1817
Families

Ursoidea is a superfamily of arctoid carnivoran mammals that includes the families Subparictidae, [1] [2] Amphicynodontidae, [3] [2] and Ursidae. The last family includes the extant lineages of bears, as well as the extinct Hemicyoninae [4] [5] and Ursavinae. [6]

The interrelationships of ursoids has had slight arrangements. In the past it was thought the extinct Amphicyonidae were stem-bears based on morphological analysis of the ear region, [7] though the most recent publications on early amphicyonids suggests they were basal caniforms. [8] [9] [10]

The amphicynodontids are sometimes classified as either a subfamily of bears, [3] a paraphyletic assemblage of early bears, [2] or even stem-pinnipeds. [11] [12] [13] The subparictids were previously classified as amphicynodontine/ids. [2] The hemicyonines have been occasionally reclassified as a separate family. [14]

Related Research Articles

<span class="mw-page-title-main">Bear</span> Family of carnivoran mammals

Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout most of the Northern Hemisphere and partially in the Southern Hemisphere. Bears are found on the continents of North America, South America, and Eurasia. Common characteristics of modern bears include large bodies with stocky legs, long snouts, small rounded ears, shaggy hair, plantigrade paws with five nonretractile claws, and short tails.

<span class="mw-page-title-main">Carnivora</span> Order of mammals

Carnivora is an order of placental mammals that have specialized in primarily eating flesh, whose members are formally referred to as carnivorans. The order Carnivora is the fifth largest order of mammals, comprising at least 279 species.

<span class="mw-page-title-main">Amphicyonidae</span> Extinct family of carnivores

Amphicyonidae is an extinct family of terrestrial carnivorans belonging to the suborder Caniformia. They first appeared in North America in the middle Eocene, spread to Europe by the late Eocene, and further spread to Asia and Africa by the early Miocene. They had largely disappeared worldwide by the late Miocene, with the latest recorded species at the end of the Miocene in Africa. They were among the first carnivorans to evolve large body size. Amphicyonids are colloquially referred to as "bear-dogs".

<span class="mw-page-title-main">Caniformia</span> Suborder of mammals

Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs, bears, raccoons, and mustelids. The Pinnipedia are also assigned to this group. The center of diversification for the Caniformia is North America and northern Eurasia. Caniformia stands in contrast to the other suborder of Carnivora, the Feliformia, the center of diversification of which was in Africa and southern Asia.

<i>Amphicyon</i> Extinct genus of carnivores

Amphicyon is an extinct genus of large carnivorans belonging to the family Amphicyonidae, subfamily Amphicyoninae, from the Miocene epoch. Members of this family received their vernacular name for possessing bear-like and dog-like features. They ranged over North America, Europe, Asia, and Africa.

<span class="mw-page-title-main">John Day Formation</span>

The John Day Formation is a series of rock strata exposed in the Picture Gorge district of the John Day River basin and elsewhere in north-central Oregon in the United States. The Picture Gorge exposure lies east of the Blue Mountain uplift, which cuts southwest–northeast through the Horse Heaven mining district northeast of Madras. Aside from the Picture Gorge district, which defines the type, the formation is visible on the surface in two other areas: another exposure is in the Warm Springs district west of the uplift, between it and the Cascade Range, and the third is along the south side of the Ochoco Mountains. All three exposures, consisting mainly of tuffaceous sediments and pyroclastic rock rich in silica, lie unconformably between the older rocks of the Clarno Formation below and Columbia River basalts above.

<i>Simocyon</i> Extinct genus of carnivores

Simocyon is a genus of extinct carnivoran mammal in the family Ailuridae. Simocyon, which was about the size of a mountain lion, lived in the late Miocene and early Pliocene epochs, and has been found in Europe, Asia, and rarely, North America and Africa.

Parictis is an extinct arctoid belonging to the family Subparictidae.

<i>Daphoenodon</i> Extinct genus of carnivores

Daphoenodon is an extinct genus of terrestrial carnivore, which lived in the early Miocene and belonged to the family Amphicyonidae of the suborder Caniformia. The species of Daphoenodon are characterized by limbs that are specialized in fore and aft movement, as well as a body alignment that results in a lengthened stride.

<i>Ysengrinia</i> Extinct genus of mammals known as bear dogs

Ysengrinia is an extinct genus of carnivoran in the family Amphicyonidae (beardogs), which lived in Europe, Asia, and North America during the Early Miocene. It was also reported from Egypt and Namibia, but this material has been reassigned to other genera of beardogs.

<span class="mw-page-title-main">Arctoidea</span> Infraorder of mammals

Arctoidea is a clade of mostly carnivorous mammals which include the extinct Hemicyonidae (dog-bears), and the extant Musteloidea, Pinnipedia, and Ursidae (bears), found in all continents from the Eocene, 46 million years ago, to the present. The oldest group of the clade is the bears, as their CMAH gene is still intact. The gene became non-functional in the common ancestor of the Mustelida. Arctoids are caniforms, along with dogs (canids) and extinct bear dogs (Amphicyonidae). The earliest caniforms were superficially similar to martens, which are tree-dwelling mustelids. Together with feliforms, caniforms compose the order Carnivora; sometimes Arctoidea can be considered a separate suborder from Caniformia and a sister taxon to Feliformia.

<span class="mw-page-title-main">Hemicyoninae</span> Extinct subfamily of bears

Hemicyoninae is an extinct subfamily of Ursidae, often called dog bears. They were bear-like carnivorans living in Europe, North America, Africa and Asia during the Oligocene through Miocene epochs 33.9–5.3 Ma, existing for approximately 28.6 million years. They are sometimes classified as a separate family.

<i>Palaeogale</i> Extinct genus of carnivores

Palaeogale is an extinct genus of carnivorous mammal known from the Late Eocene, Oligocene, and Early Miocene of North America, Europe, and Eastern Asia. A small carnivore often associated with the mustelids, Palaeogale might have been similar to living genets, civets, and linsangs.

<i>Gustafsonia</i> Extinct genus of carnivores

Gustafsonia is an extinct genus of carnivoran belonging to the family Amphicyonidae. The type species, Gustafsonia cognita, was described in 1986 by Eric Paul Gustafson, who originally interpreted it as a miacid and named it Miacis cognitus. It was subsequently considered to be the only species of the diverse genus Miacis that belonged to the crown-group Carnivora, within the Caniformia, and it was ultimately assigned to the family Amphicyonidae. The type specimen or holotype was discovered in Reeve's bonebed, western Texas, in the Chambers Tuff Formation in 1986. The University of Texas holds this specimen. It is the only confirmed fossil of this species.

<i>Brachyrhynchocyon</i> Extinct genus of carnivores

Brachyrhynchocyon is an extinct genus of terrestrial carnivore, which belonged to the family Amphicyonidae of the suborder Caniformia.

The Sharps Formation is a geologic formation in South Dakota. It preserves fossils dating back to the Paleogene.

<span class="mw-page-title-main">Amphicynodontidae</span> Extinct clade of mammals

Amphicynodontidae is a probable clade of extinct arctoids. While some researchers consider this group to be an extinct subfamily of bears, a variety of morphological evidence links amphicynodontines with pinnipeds, as the group were semi-aquatic otter-like mammals. In addition to the support of the pinniped–amphicynodontine clade, other morphological and some molecular analyses support bears being the closest living relatives to pinnipeds. According to McKenna and Bell (1997) Amphicynodontinae are classified as stem-pinnipeds in the superfamily Phocoidea. Fossils of these mammals have been found in Europe, North America and Asia. Amphicynodontines should not be confused with Amphicyonids (bear-dogs), a separate family of Carnivora which is a sister clade to arctoids within the caniforms, but which may be listed as a clade of extinct arctoids in older publications.

Namibiocyon is an extinct genus of carnivoran mammals, belonging to the family Amphicyonidae, that lived in Namibia during the Early Miocene epoch. Before the erection of this taxon in 2022, the type and only species, N. ginsburgi, had been assigned to a variety of other genera. It is notable for its adaptions toward hypercarnivory.

Subparictidae is an extinct family of early Paleogene arctoid carnivorans endemic to North America that closely related to bears. They were small, raccoon-like mammals that lived from the Eocene to the early Miocene. This family includes a handful of genera such as Subparictis, Parictis, Nothocyon, and Eoarctos.

<i>Amphicticeps</i> Extinct genus of carnivores

Amphicticeps is an extinct genus of small, weasel-like carnivoran mammal. It lived in Mongolia during the Oligocene. The genus was erected in 1924 for the species A. shackelfordi on the basis of a well-preserved skull. Historically, the systematic position of this genus has been problematic until more specimens were described decades later.

References

  1. Baskin, J. A.; Tedford, R. H. (1996). Small arctoid and feliform carnivorans. pp. 486–497. doi:10.1017/CBO9780511665431.025. ISBN   978-0-521-43387-7.{{cite book}}: |journal= ignored (help)
  2. 1 2 3 4 Wang, Xiaoming; Emry, Robert J.; Boyd, Clint A.; Person, Jeff J.; White, Stuart C.; Tedford, Richard H. (2022). "An exquisitely preserved skeleton of Eoarctos vorax (Nov. Gen. Et sp.) from Fitterer Ranch, North Dakota (Early Oligocene) and systematics and phylogeny of North American early arctoids (Carnivora, Caniformia)". Journal of Vertebrate Paleontology. 42: 1–123. doi: 10.1080/02724634.2022.2145900 . S2CID   259025727.
  3. 1 2 McLellan, B.; Reiner, D.C. (1992). "A review of bear evolution" (PDF). International Association for Bear Research and Management. 9 (1): 85–96. doi:10.2307/3872687. JSTOR   3872687. S2CID   91124592. Archived from the original (PDF) on 2020-02-19.
  4. Louis De Bonis (2013). "Ursidae (Mammalia, Carnivora) from the Late Oligocene of the "Phosphorites du Quercy" (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862". Geodiversitas. 35 (4): 787–814. doi:10.5252/g2013n4a4. S2CID   131561629.
  5. L. de Bonis (2011). "A new species of Adelpharctos (Mammalia, Carnivora, Ursidae) from the late Oligocene of the "Phosphorites du Quercy" (France)". Estudios Geológicos. 67 (2): 179–186. doi: 10.3989/egeol.40553.181 .
  6. Qiu, Zhan-Xiang; et al. (2014). "A Late Miocene Ursavus skull from Guanghe, Gansu, China". Vertebrata PalAsiatica. 52 (3): 265–302.
  7. Hunt Jr., Robert M. (2001). "Small Oligocene Amphicyonids from North America (Paradaphoenus, Mammalia, Carnivora)". American Museum Novitates (3331): 1-20. doi:10.1206/0003-0082(2001)331<0001:SOAFNA>2.0.CO;2. ISSN   0003-0082. S2CID   198160461.
  8. Hunt, Robert M. Jr. (2004). "Global Climate and the Evolution of Large Mammalian Carnivores during the Later Cenozoic in North America" (PDF). Cenozoic Carnivores and Global Climate. pp. 139–285. doi:10.1206/0003-0090(2004)285<0139:C>2.0.CO;2. S2CID   86236545. Archived from the original (PDF) on July 20, 2007.{{cite book}}: |journal= ignored (help)
  9. Morlo, Michael; Miller, Ellen R.; El-Barkooky, Ahmed N. (2007). "Creodonta and Carnivora from Wadi Moghra, Egypt". Journal of Vertebrate Paleontology. 27 (1): 145-159. doi:10.1671/0272-4634(2007)27[145:CACFWM]2.0.CO;2. ISSN   0272-4634. S2CID   86235694.
  10. Tomiya, Susumu; Tseng, Zhijie Jack (2016). "Whence the beardogs? Reappraisal of the Middle to Late Eocene 'Miacis' from Texas, USA, and the origin of Amphicyonidae (Mammalia, Carnivora)". Royal Society Open Science. 3 (10): 160518. Bibcode:2016RSOS....360518T. doi:10.1098/rsos.160518. PMC   5098994 . PMID   27853569.
  11. Tedford, R. H.; Barnes, L. G.; Ray, C. E. (1994). "The early Miocene littoral ursoid carnivoran Kolponomos: Systematics and mode of life" (PDF). Proceedings of the San Diego Society of Natural History. 29: 11–32. Archived from the original (PDF) on 2012-09-22. Retrieved 24 July 2010.
  12. Rybczynski, N.; Dawson, M.R.; Tedford, R.H. (2009). "A semi-aquatic Arctic mammalian carnivore from the Miocene epoch and origin of Pinnipedia". Nature. 458 (7241): 1021–24. Bibcode:2009Natur.458.1021R. doi:10.1038/nature07985. PMID   19396145. S2CID   4371413.
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  14. McKenna, Malcolm C., and Bell, Susan K. 1997. Classification of Mammals Above the Species Level. Columbia University Press, New York, 631 pp.  ISBN   0-231-11013-8