| Arabosteus Temporal range: | |
|---|---|
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Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | † Placodermi |
| Order: | † Acanthothoraci |
| Genus: | † Arabosteus Olive et al., 2011 |
| Species: | †A. variabilis |
| Binomial name | |
| †Arabosteus variabilis Olive et al., 2011 | |
Arabosteus is an extinct genus of palaecanthaspid placoderm that inhabited what is now Saudi Arabia during the Early Devonian epoch. Its only species is Arabosteus variabilis. [1]
Placoderms are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates, and the rest of the body was scaled or naked depending on the species.
Arthrodira is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of placoderms.
Bothriolepis was a widespread, abundant and diverse genus of antiarch placoderms that lived during the Middle to Late Devonian period of the Paleozoic Era. Historically, Bothriolepis resided in an array of paleo-environments spread across every paleocontinent, including near shore marine and freshwater settings. Most species of Bothriolepis were characterized as relatively small, benthic, freshwater detritivores, averaging around 30 centimetres (12 in) in length. However, the largest species, B. rex, had an estimated bodylength of 170 centimetres (67 in). Although expansive with over 60 species found worldwide, comparatively Bothriolepis is not unusually more diverse than most modern bottom dwelling species around today.
Rhenanida is an order of scaly placoderms. Unlike most other placoderms, the rhenanids' armor was made up of a mosaic of unfused scales and tubercles. The patterns and components of this "mosaic" correspond to the plates of armor in other, more advanced placoderms, suggesting that the ancestral placoderm had armor made of unfused components, as well.
Phyllolepida is an extinct taxon of flattened placoderms found throughout the world, with fossils being found in Devonian strata. Like other flattened placoderms, the phyllolepids were bottom-dwelling predators that ambushed prey. Unlike other flattened placoderms, the phyllolepids were inhabitants of freshwater environments.
Acanthothoraci is an extinct group of chimaera-like placoderms closely related to the rhenanid placoderms. Superficially, the acanthoracids resembled scaly chimaeras and (relatively) heavily armored ptyctodonts. They were distinguished from chimaeras by their large scales and plates, a pair of large spines that emanate from their chests, tooth-like beak plates, and the typical bone-enhanced placoderm eyeball. They were distinguished from other placoderms by differences in skull anatomy and by patterns on the skull plates and thoracic plates that are unique to this order.
Brindabellaspis stensioi is a placoderm with a flat, platypus-like snout from the Early Devonian of the Taemas-Wee Jasper reef in Australia. When it was first discovered in 1980, it was originally regarded as a Weejasperaspid acanthothoracid due to anatomical similarities with the other species found at the reef.
Phyllolepis is the type genus of Phyllolepida, an extinct taxon of arthrodire placoderm fish from the middle to late Devonian. The species of Phyllolepis, themselves, are restricted to the Famennian-aged freshwater strata of the Late Devonian, around 360 million years ago. Fossils of this genus have been found primarily in Europe and North America. The end of the Devonian saw them disappear in a mass extinction.
Murrindalaspis is an extinct genus of acanthothoracid placoderm found in the McLarty Member of the Murrindal Limestone, of the Early Devonian-aged Buchan Group in eastern Victoria, Australia. Murrindalaspis differs from other acanthothoracids by having a dorsal plate with a large, blade-like flattened, recurved crest emanating from the medial line, and no ventral keel. So far, the genus is known only from dorsal plates and ossified eyeballs. The genus differs from the closely related Weejasperaspis in that the dorsal crest of the latter is shorter, and triangular-shaped.
Austrophyllolepis is an extinct genus of phyllolepid arthrodire placoderm from Middle to Late Devonian freshwater strata of Australia. The type species, A. ritchiei is found in Givetian to early Frasnian-aged freshwater strata near what is now Mount Howitt. A second species, A. dulciensis, is found from Middle Devonian freshwater strata from the Dulcie Sandstone of Georgina Basin, Central Australia.
Fallacosteus is an extinct monospecific genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period, found at the Gogo Formation of Kimberley, Western Australia. As with almost all other camuropiscids, F. turneri had an elongated snout that may have enhanced its hydrodynamic streamlining.
Nefudina qalibahensis is a rhenanid placoderm from the early Emsian during the Early Devonian epoch of Northeastern Saudi Arabia. It is known from skull fragments and scales.
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.
Homostiidae is a family of flattened arthrodire placoderms from the Early to Middle Devonian. Fossils appear in various strata in Europe, Russia, Morocco, Australia, Canada and Greenland.
Palaeacanthaspis vasta is an extinct acanthothoracid placoderm from the early Devonian of Europe.
Simblaspis is an extinct monospecific genus of arthrodire placoderm fish of the Early Devonian period. The type species Aleosteus eganensis was described in 1958, and was found in Pragian strata of the Qasr Limestone in Saudi Arabia.
Actinolepis is an extinct genus of actinolepid placoderm from the Early Devonian. Four species are known: A. magna from Estonia, A. spinosa from Latvia, the type species A. tuberculata from New Zealand and A. zaikai from Belarus.
Kujdanowiaspis is an extinct genus of arthrodire placoderm fish from the Early Devonian of Podolia (Ukraine), Poland and Spain. Kujdanowiaspis is known from many fragmentary head shields and body armours.
Millerosteus is an extinct genus of coccosteid arthrodire placoderm from the Early Givetian stage of the Middle Devonian period. Fossils are found in Orkney and Caithness, Scotland. It was a small placoderm with a body length of 14 cm (5.5 in). Millerosteus is one of the few arthrodires known from specimens preserving the entire skeleton.
Dickosteus is an extinct genus of coccosteid arthrodire placoderm from the Late Eifelian to Early Givetian stages of the Middle Devonian period. Fossils are found in Orkney and Caithness, Scotland. It was a small placoderm with a total body length of 43.7 cm (17.2 in). It is one of the few placoderms for which complete bodies are known.
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