Australaves | |
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Kestrel, Falco tinnunculus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Clade: | Telluraves |
Clade: | Australaves Ericson, 2012 |
Clades | |
Australaves is a clade of birds, [3] defined in 2012, [4] consisting of the Eufalconimorphae (passerines, parrots and falcons) as well as the Cariamiformes (including seriemas and the extinct "terror birds"). [5] They appear to be the sister group of Afroaves. [5]
The clade's name, meaning 'southern birds', reflects the group's evolutionary origins in the Southern Hemisphere: passerines and parrots in Australia, and falcons and sereimas in South America. [4]
As in the case of Afroaves, the most basal clades have predatory extant members, suggesting this was the ancestral lifestyle; [6] however, some researchers like Darren Naish are skeptical of this assessment, since some extinct representatives such as the herbivorous Strigogyps led other lifestyles. [7] Basal parrots and falcons are at any rate vaguely crow-like and probably omnivorous. [8]
Australaves |
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Cladogram of Telluraves relationships based on Kuhl et al. (2020) and Braun & Kimball (2021) [2] [9]
Near passerines and higher land-bird assemblage are terms of traditional, pre-cladistic taxonomy that have often been given to tree-dwelling birds or those most often believed to be related to the true passerines owing to morphological and ecological similarities; the group corresponds to some extent with the Anomalogonatae of Alfred Henry Garrod.
The Pelecaniformes are an order of medium-sized and large waterbirds found worldwide. As traditionally—but erroneously—defined, they encompass all birds that have feet with all four toes webbed. Hence, they were formerly also known by such names as totipalmates or steganopodes. Most have a bare throat patch, and the nostrils have evolved into dysfunctional slits, forcing them to breathe through their mouths. They also have a pectinate nail on their longest toe. This is shaped like a comb and is used to brush out and separate their feathers. They feed on fish, squid, or similar marine life. Nesting is colonial, but individual birds are monogamous. The young are altricial, hatching from the egg helpless and naked in most. They lack a brood patch.
The mesites (Mesitornithidae) are a family of birds that are part of a clade (Columbimorphae) that include Columbiformes and Pterocliformes. They are smallish flightless or near flightless birds endemic to Madagascar. They are the only family with more than two species in which every species is threatened.
Neoaves is a clade that consists of all modern birds with the exception of Palaeognathae and Galloanserae. Almost 95% of the roughly 10,000 known species of extant birds belong to the Neoaves.
Apodimorphae is a clade of strisorean birds that include the extant families Trochilidae (hummingbirds), Hemiprocnidae (treeswifts), Apodidae (swifts), and Aegothelidae (owlet-nightjars), as well as many fossil families. This grouping of birds has been supported in a variety of recent studies. There are two higher classification schemes that have been proposed for the apodimorph families. One is all strisorean birds are classified in the order Caprimulgiformes, while the other is the strisorean birds are split into several distinct orders. In this case Apodimorphae is a subclade of Strisores that includes the orders Aegotheliformes and the Apodiformes. A similar name for the group Daedalornithes has been used for the owlet-night-apodiform clade, there is a difference between the two names with Apodimorphae defined as the total-group and Daedalornithes defined as the crown group.
Mirandornithes is a clade that consists of flamingos and grebes. Many scholars use the term Phoenicopterimorphae for the superorder containing flamingoes and grebes.
Aequornithes, or core water birds, are defined as "the least inclusive clade containing Gaviidae and Phalacrocoracidae".
The order Falconiformes is represented by the extant family Falconidae and a handful of enigmatic Paleogene species. Traditionally, the other bird of prey families Cathartidae, Sagittariidae (secretarybird), Pandionidae (ospreys), Accipitridae (hawks) were classified in Falconiformes. A variety of comparative genome analyses published since 2008, however, found that falcons are part of a clade of birds called Australaves, which also includes seriemas, parrots and passerines. Within Australaves falcons are more closely related to the parrot-passerine clade than they are to the seriemas. The hawks, vultures and owls are placed in the clade Afroaves.
Psittacopasseres is a taxon of birds consisting of the Passeriformes and Psittaciformes (parrots). Per Ericson and colleagues, in analysing genomic DNA, revealed a lineage comprising passerines, psittacines and Falconiformes. The group was proposed following an alignment of nuclear intron sequences by Shannon Hackett et al. in 2008. It was formally named as Psittacopasserae in a 2011 Nature Communications article by Alexander Suh and other authors working with Jürgen Schmitz's group, based on genetic analysis of the insertion of retroposons into the genomes of key avian lineages over the course of evolution during the Mesozoic Era.
Cariamiformes is an order of primarily flightless birds that has existed for over 50 million years. The group includes the family Cariamidae (seriemas) and the extinct families such as Phorusrhacidae, Bathornithidae, Idiornithidae and Ameghinornithidae. Extant members (seriemas) are only known from South America, but fossils of many extinct taxa are also found in other continents including Europe and North America. Though traditionally considered a suborder within Gruiformes, both morphological and genetic studies show that it belongs to a separate group of birds, Australaves, whose other living members are Falconidae, Psittaciformes and Passeriformes.
Eufalconimorphae is a proposed clade of birds, consisting of passerines, parrots, falcons, caracaras, and forest falcons. It has whole-genome DNA support. Eufalconimorphae birds are characterized by their strong and hooked beaks, sharp talons, and powerful wings. They have excellent eyesight, which allows them to spot their prey from great distances. The Eufalconimorphae is noted to produce aerodynamic force during the upstroke of flight to help create a vertical flight pattern.
Strisores, sometimes called nightbirds, is a clade of birds that includes the living families and orders Caprimulgidae, Nyctibiidae (potoos), Steatornithidae (oilbirds), Podargidae (frogmouths), Apodiformes, as well as the Aegotheliformes (owlet-nightjars) whose distinctness was only recently realized. The Apodiformes and the Aegotheliformes form the Daedalornithes.
Afroaves is a clade of birds, consisting of the kingfishers and kin (Coraciiformes), woodpeckers and kin (Piciformes), hornbills and kin (Bucerotiformes), trogons (Trogoniformes), cuckoo roller (Leptosomiformes), mousebirds (Coliiformes), owls (Strigiformes), raptors (Accipitriformes) and New World vultures (Cathartiformes). The most basal clades are predatory, suggesting the last common ancestor of Afroaves was also a predatory bird.
Telluraves is a recently defined clade of birds defined by their arboreality. Based on most recent genetic studies, the clade unites a variety of bird groups, including the australavians as well as the afroavians. They appear to be the sister group of the Phaethoquornithes.
Passerea is a clade of neoavian birds that was proposed by Jarvis et al. (2014). Their genomic analysis recovered two major clades within Neoaves, Passerea and Columbea, and concluded that both clades appear to have many ecologically driven convergent traits.
Gruimorphae is a clade of birds that contains the orders Charadriiformes and Gruiformes identified by molecular analysis. This grouping has had historical support, as various charadriiform families such as the families Pedionomidae and Turnicidae were classified as gruiforms. It may also have support from the fossil record since the discovery of Nahmavis from the Early Eocene of North America.
Phaethoquornithes is a clade of birds that contains Eurypygimorphae and Aequornithes, which was first recovered by genome analysis in 2014. Members of Eurypygimorphae were originally classified in the obsolete group Metaves, and Aequornithes were classified as the sister taxon to Musophagiformes or Gruiformes.
Columbimorphae is a clade discovered by genome analysis that includes birds of the orders Columbiformes, Pterocliformes (sandgrouse), and Mesitornithiformes (mesites). Previous analyses had also recovered this grouping, although the exact relationships differed. Some studies indicated a sister relationship between sandgrouse and pigeons while other studies favored a sister grouping of mesites and sandgrouse instead.
Vanescaves is a probable clade of strisorean birds that include the clades Steatornithiformes, Nyctibiiformes, Podargiformes (frogmouths), and Apodimorphae. Some molecular studies do support the grouping of these birds, others offer conflicting positions of the non-apodimorphaean strisoreans. In 2019 the authors Chen et al. performed a combined analysis using 2289 ultra-conserved elements [UCEs], 117 morphological characters from extant and fossil taxa found support in this clade. The authors then proposed to name this group, which its meaning is Latin for "vanish birds" in reference to the disparate nature of their geographic distribution, as well as to the poem "A Route of Evanescence" by the American poet Emily Dickinson which features a hummingbird as the main subject. In 2020 Chen & Field named the two major subclades of this group, with Sedentaves and Letornithes for their crown-groups.
Litusilvanae is a proposed clade of birds, position as the sister clade to Aequorlitornithes. This clade comprises Gruimorphae and Strisores. While different lines of evidence from molecular, morphology and the fossil record has found support in the clades Gruimorphae and Strisores Wu et al. (2024) was the first to find support in such a novel sister group relationship between these two taxa.