Ophioglossaceae, the adder's-tongue family, is a small family of ferns. In the Pteridophyte Phylogeny Group classification of 2016, it is the only family in the order Ophioglossales, which together with the Psilotales is placed in the subclass Ophioglossidae. The Ophioglossidae are one of the groups traditionally known as eusporangiate ferns. Members of the family differ from other ferns in a number of ways. Many have only a single fleshy leaf at a time. Their gametophytes are subterranean and rely on fungi for energy.
Botrychium is a genus of ferns, seedless vascular plants in the family Ophioglossaceae. Botrychium species are known as moonworts. They are small, with fleshy roots, and reproduce by spores shed into the air. One part of the leaf, the trophophore, is sterile and fernlike; the other, the sporophore, is fertile and carries the clusters of sporangia or spore cases. Some species only occasionally emerge above ground and gain most of their nourishment from an association with mycorrhizal fungi.
Asplenium platyneuron, commonly known as ebony spleenwort or brownstem spleenwort, is a fern native to North America east of the Rocky Mountains. It takes its common name from its dark, reddish-brown, glossy stipe and rachis, which support a once-divided, pinnate leaf. The fertile fronds, which die off in the winter, are darker green and stand upright, while the sterile fronds are evergreen and lie flat on the ground. An auricle at the base of each pinna points towards the tip of the frond. The dimorphic fronds and alternate, rather than opposite, pinnae distinguish it from the similar black-stemmed spleenwort.
Sceptridium is a genus of seedless vascular plants in the family Ophioglossaceae, closely allied to the genus Botrychium. It is also closely related to the genus Botrypus. Sceptridium species are commonly called the grape-ferns.
Botrypus virginianus, synonym Botrychium virginianum, sometimes called rattlesnake fern is a species of perennial fern in the adders-tongue family. It is monotypic within the genus Botrypus, meaning that it is the only species within the genus. It is called the rattlesnake fern in some parts of North America, due to its habit of growing in places where rattlesnakes are also found. Rattlesnake fern prefers to grow in rich, moist woods in dense shade and will not tolerate direct sunlight.
Botrychium pumicola, with the common name pumice moonwort, is a rare fern.
Sceptridium multifidum is a fern species in the Ophioglossaceae, known by the common names leathery grapefern and leathery moonwort.
Botrychium ascendens is a species of fern in the family Ophioglossaceae known by the common names triangle-lobe moonwort and upswept moonwort. It is native to North America from British Columbia to northern California as well as parts of eastern Canada. It lives in different habitat types, including grassy riverside areas. This is very small plant growing from an underground caudex and sending one yellow-green leaf above the surface of the ground. The leaf is up to 6 centimeters tall and is divided into a sterile and a fertile part. The sterile part of the leaf has fan-shaped or wedge-shaped leaflets. The fertile part of the leaf is very different in shape, with tiny grapelike clusters of sporangia by which it reproduces.
Botrychium crenulatum is a species of fern in the family Ophioglossaceae known by the common names scalloped moonwort and dainty moonwort. It is native to North America from British Columbia to California to Wyoming, where it is uncommon throughout most of its range, appearing incidentally at scattered spots on wet meadows in coniferous forests and marshy areas such as swamps. This is very small plant growing from an underground caudex and sending one thin, shiny, yellow-green leaf above the surface of the ground. The leaf is up to about 6 centimeters tall and is divided into a sterile and a fertile part. The sterile part of the leaf has veined, fan-shaped leaflets with wrinkly edges. The fertile part of the leaf is very different in shape, with tiny grapelike clusters of sporangia by which it reproduces.
Botrychium lunaria is a species of fern in the family Ophioglossaceae known by the common name moonwort or common moonwort. It is the most widely distributed moonwort, growing throughout the Northern Hemisphere across Eurasia and from Alaska to Greenland, as well as temperate parts of the Southern Hemisphere.
Botrychium montanum is a species of fern in the family Ophioglossaceae known by the common names western goblin and mountain moonwort. It is native to western North America from British Columbia to northern California to Montana, where it grows in the dark understory of coniferous forests and other moist wooded areas. This is very small plant growing from an underground caudex and sending one thin gray-green leaf above the surface of the ground. The leaf is less than 8 centimeters tall and is divided into a sterile and a fertile part. The sterile part of the leaf has irregularly shaped angled leaflets. The fertile part of the leaf is very different in shape, with grapelike clusters of sporangia by which it reproduces.
Botrychium pinnatum is a species of fern in the family Ophioglossaceae, known by the common name northwestern moonwort. It is native to North America from Alaska to northern Canada to California and Arizona, where it is generally scattered and uncommon, growing in coniferous forests and grassy meadows. This is very small plant growing from an underground caudex and sending one thin, shiny, green leaf above the surface of the ground. The leaf is less than 8 centimeters tall and is divided into a sterile and a fertile part. The flat sterile part of the leaf has oval to widely lance-shaped leaflets. The fertile part of the leaf is very different in shape, with grapelike clusters of sporangia by which it reproduces.
Sceptridium dissectum is a common fern in the family Ophioglossaceae, occurring in eastern North America. Like other plants in this group, it normally only sends up one frond per year. It has long been the subject of confusion because the frond presents in one of two forms, either the normal form that resembles other plants in the genus, or the skeletonized form.
Botrychium matricariifolium is a species of fern in the Ophioglossaceae family. It is referred to by the common names chamomile grape-fern, daisyleaf grape-fern, and matricary grape-fern. It is native to Europe and parts of eastern North America, including eastern Canada and parts of the United States.
Botrychium paradoxum is a species of fern in the family Ophioglossaceae known by the common name peculiar moonwort. It is native to North America, where there are scattered occurrences in Alberta, British Columbia, Saskatchewan, Montana, Oregon, Utah, Washington, and Wyoming.
Asplenium resiliens, the blackstem spleenwort or little ebony spleenwort, is a species of fern native to the Western Hemisphere, ranging from the southern United States south to Uruguay, including parts of the Caribbean. Found on limestone substrates, it is named for its distinctive purplish-black stipe and rachis. A triploid, it is incapable of sexual reproduction and produces spores apogamously. First described by Martens and Galeotti in 1842 under the previously used name Asplenium parvulum, the species was given its current, valid name by Kunze in 1844. Several similar species are known from the tropics; A. resiliens may have arisen from these species by reticulate evolution, but precise relationships among the group are not yet certain.
Asplenium tutwilerae is a rare epipetric fern found only in Hale County, Alabama, United States. A. tutwilerae is a fertile allotetraploid, formed by the chromosomal doubling of a specimen of the sterile diploid A. × ebenoides, a hybrid of A. platyneuron and A. rhizophyllum. Except for its spores, which are fertile rather than malformed, A. tutwilerae is essentially identical to A. × ebenoides and was described as part of that species until 2007. It is named in honor of Julia Tutwiler, who discovered the only known wild population at Havana Glen in 1873.
Sceptridium oneidense, the blunt-lobed grapefern, is a fern species in the family Ophioglossaceae.
Anemia tabascana is a fern endemic to the state of Tabasco, Mexico. It grows on road banks on a single hill in Huimanguillo. Like other members of the genus, its leaves are pinnately divided into a single set of leaflets; in fertile leaves, the lowest pair of leaflets projects at right angles to the rest of the leaf and bears spores. The freely forking veins, which do not rejoin one another after forking, and sparse, rather than abundant, hairs on the leaf axes, distinguish it from similar species in the genus.
Botrychium campestre is a fern species in Ophioglossaceae, commonly called prairiemoonwort, prairie dunewort, Iowa moonwort, or plains grapefern. It was first discovered in 1982 and described a few years later.
