Harpaphe haydeniana, commonly known as the yellow-spotted millipede, almond-scented millipede or cyanide millipede, is a species of polydesmidan ("flat-backed") millipede found in the moist forests along the Pacific coast of North America, from Southeast Alaska to California. The dark coloration with contrasting yellow-tipped keels warn of its ability to exude toxic hydrogen cyanide as a defense. Despite the various common names given the species, the coloration pattern, cyanide defense, and associated almond scent occur in other flat-backed millipedes around the world.
Polydesmida is the largest order of millipedes, containing approximately 3,500 species, including all the millipedes reported to produce hydrogen cyanide (HCN).
William Tinsley Keeton was an American zoologist known internationally for his work on animal behavior, especially bird migration, and for his work on millipede taxonomy. He was a well-liked professor of biology at Cornell University in Ithaca, New York and author of a widely used introductory textbook, Biological Science.
Xystodesmidae is a family of millipedes. Its members often have very small distributional areas, with many species only known from a single locality. They are found across the northern hemisphere, with peak diversity in the Appalachian Mountains, where one-third of the 300 or so species occur. They are particularly abundant in deciduous broadleaf forests in the Mediterranean Basin, Africa, Asia, Central and North America, and Russia. Information on basic taxonomy is scant for this family; for example, it is estimated that the genus Nannaria contains over 200 species, but only 25 have so far been described.
Motyxia is a genus of cyanide-producing millipedes that are endemic to the southern Sierra Nevada, Tehachapi, and Santa Monica mountain ranges of California. Motyxias are blind and produce the poison cyanide, like all members of the Polydesmida. All species have the ability to glow brightly: some of the few known instances of bioluminescence in millipedes.
Richard Lawrence Hoffman was an American zoologist known as an international expert on millipedes, and a leading authority on the natural history of Virginia and the Appalachian Mountains. He was a biology professor at Virginia's Radford College for almost thirty years, and curator of invertebrates at the Virginia Museum of Natural History for another twenty years. He co-founded the Virginia Natural History Society, described over 400 species of millipedes, and produced more than 480 scientific publications. He is commemorated in the scientific and/or common names of over 30 animal species, including the valley and ridge salamander and Hoffman's dwarf centipede.
Brachoria is a genus of polydesmidan millipedes in the family Xystodesmidae inhabiting the Eastern United States. Also known as the Appalachian mimic millipedes, at least 30 species are known, with highest diversity in the Appalachian Mountains, especially the Cumberland Plateau and Ridge and Valley Province.
Xystocheir is a genus of millipedes in the family Xystodesmidae. The genus is endemic to California in the United States, where it is distributed in the Coast Ranges and the Sierra Nevada.
Parafontaria is a genus of "flat-backed" millipedes consisting of 13 species native to Japan, where they are referred to as train millipedes. This is because some species exhibit periodical swarming behavior during which large numbers congregate and can impact train passage when this congregation occurs on tracks. Documentation of this event goes back to 1920. Individuals vary from around 3.5 to 6 cm as adults, and feed on leaf litter as well as soil, making them comparable to earthworms in facilitating decomposition and soil nutrient cycling.
Apheloria tigana, Yellow-and-black Flat Millipede, is a large North American flat-backed millipede in the family Xystodesmidae. It is reported to secrete cyanide compounds as a defense. It is recommended that one wash hands after handling this organism as the toxic compounds it secretes are poisonous and can cause extreme irritation if rubbed in the eyes.
Apheloria is a genus of flat-backed millipedes in the family Xystodesmidae, occurring in the central and southeastern United States, and ranging as far north as southern Quebec, Canada. They are aposematically colored in black and contrasting reds and yellows, and some species in the Appalachian Mountains resemble species of Brachoria where they co-occur, a phenomenon known as Müllerian mimicry.
Pachydesmus is a genus of flat-backed millipedes in the family Xystodesmidae. There are about 12 described species in Pachydesmus.
Semionellus is a genus of flat-backed millipedes in the family Xystodesmidae. There are at least three described species in Semionellus.
Selenocheir is a genus of flat-backed millipedes in the family Xystodesmidae. There are at least three described species in Selenocheir.
Thrinaxoria is a genus of flat-backed millipedes in the family Xystodesmidae. There are at least three described species in Thrinaxoria.
Boraria is a genus of flat-backed millipedes in the family Xystodesmidae. There are about seven described species in Boraria.
Montaphe is a genus of flat-backed millipedes in the family Xystodesmidae. There are at least two described species in Montaphe.
Furcillaria is a genus of flat-backed millipedes in the family Xystodesmidae. There are at least four described species in Furcillaria.
Cherokia is a genus of flat-backed millipedes in the family Xystodesmidae. There are at least three described species in Cherokia.
