Columbea Temporal range: [1] | |
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Victoria crowned pigeon ( Goura victoria ) | |
Black-necked Grebe ( Podiceps nigricollis ) | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Infraclass: | Neognathae |
Clade: | Neoaves |
Clade: | Columbea Jarvis et al., 2014 |
Clades | |
Columbea is a clade suggested by genome analysis that contains Columbiformes (pigeons and doves), Pteroclidae (sandgrouse), Mesitornithidae (mesites) and Mirandornithes (flamingos and grebes). [2] Until their recent placement as the sister taxon to Passerea, in the last decade various genetic analysis found them to be in the obsolete clade Metaves. [3] [4]
Columbea |
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The mesites (Mesitornithidae) are a family of birds that are part of a clade (Columbimorphae) that include Columbiformes and Pterocliformes. They are smallish flightless or near flightless birds endemic to Madagascar. They are the only family with more than two species in which every species is threatened.
The Phaethontiformes are an order of birds. They contain one extant family, the tropicbirds (Phaethontidae), and one extinct family Prophaethontidae from the early Cenozoic. Several fossil genera have been described.
Neoaves is a clade that consists of all modern birds with the exception of Paleognathae and Galloanserae. Almost 95% of the roughly 10,000 known species of extant birds belong to the Neoaves.
Aequornithes, or core water birds, are defined as "the least inclusive clade containing Gaviidae and Phalacrocoracidae".
The order Falconiformes is represented by the extant family Falconidae and a handful of enigmatic Paleogene species. Traditionally, the other bird of prey families Cathartidae, Sagittariidae (secretarybird), Pandionidae (ospreys), Accipitridae (hawks) were classified in Falconiformes. A variety of comparative genome analysis published since 2008, however, found that falcons are part of a clade of birds called Australaves, which also includes seriemas, parrots and passerines. Within Australaves falcons are more closely related to the parrot-passerine clade (Psittacopasserae), which together they form the clade Eufalconimorphae. The hawks and vultures occupy a basal branch in the clade Afroaves in their own clade Accipitrimorphae, closer to owls and woodpeckers.
Cariamiformes is an order of primarily flightless birds that has existed for over 60 million years. The group includes the family Cariamidae (seriemas) and the extinct families Phorusrhacidae, Bathornithidae, Idiornithidae and Ameghinornithidae. Though traditionally considered a suborder within Gruiformes, both morphological and genetic studies show that it belongs to a separate group of birds, Australaves, whose other living members are Falconidae, Psittaciformes and Passeriformes.
Strisores, sometimes called nightbirds, is a clade of birds that includes the living families and orders Caprimulgidae, Nyctibiidae (potoos), Steatornithidae (oilbirds), Podargidae (frogmouths), Apodiformes, as well as the Aegotheliformes (owlet-nightjars) whose distinctness was only recently realized. The Apodiformes and the Aegotheliformes form the Daedalornithes.
Passerea is a clade of neoavian birds that was proposed by Jarvis et al. (2014). Their genomic analysis recovered two major clades within Neoaves, Passerea and Columbea, and concluded that both clades appear to have many ecologically driven convergent traits.
Eurypygimorphae or Phaethontimorphae is a clade of birds that contains the orders Phaethontiformes (tropicbirds) and Eurypygiformes recovered by genome analysis. The relationship was first identified in 2013 based on their nuclear genes. Historically these birds were placed at different parts of the tree, with tropicbirds in Pelecaniformes and the kagu and sunbittern in Gruiformes. Some genetic analyses have placed the eurypygimorph taxa in the controversial and obsolete clade Metaves, with uncertain placement within that group. More recent molecular studies support their grouping together in Eurypygimorphae, which is usually recovered as the sister taxon to Aequornithes within Ardeae.
Otidimorphae is a clade of birds that contains the orders Cuculiformes (cuckoos), Musophagiformes (turacos), and Otidiformes (bustards) identified in 2014 by genome analysis. While the bustards seem to be related to the turacos, other genetic studies have found the cuckoos to be closer to the bustards than the turacos are.
Phaethoquornithes is a clade of birds that contains Eurypygimorphae and Aequornithes, which was first recovered by genome analysis in 2014. Members of Eurypygimorphae were originally classified in the obsolete group Metaves, and Aequornithes were classified as the sister taxon to Musophagiformes or Gruiformes.
Austrodyptornithes is a clade of birds that include the orders Sphenisciformes (penguins) and Procellariiformes. A 2014 analysis of whole genomes of 48 representative bird species concluded that penguins are the sister group of Procellariiformes, from which they diverged about 60 million years ago.
Coraciimorphae is a clade of birds that contains the order Coliiformes (mousebirds) and the clade Cavitaves. The name however was coined in the 1990s by Sibley and Ahlquist based on their DNA-DNA hybridization studies conducted in the late 1970s and throughout the 1980s. However their Coraciimorphae only contains Trogoniformes and Coraciiformes.
Eucavitaves is a clade that contains the order Trogoniformes (trogons) and the clade Picocoraciae. The name refers to the fact that the majority of them nest in cavities.
Cavitaves is a clade that contain the order Leptosomiformes and the clade Eucavitaves. The name refers to the fact that the majority of them nest in cavities.
Picocoraciae is a clade that contains the order Bucerotiformes and the clade Picodynastornithes supported by various genetic analysis and morphological studies. While these studies supported a sister grouping of Coraciiformes and Piciformes, a large scale, sparse supermatrix has suggested alternative sister relationship between Bucerotiformes and Piciformes instead.
Columbimorphae is a clade discovered by genome analysis that includes birds of the orders Columbiformes, Pterocliformes (sandgrouse), and Mesitornithiformes (mesites). Previous analyses had also recovered this grouping, although the exact relationships differed. Some studies indicated a sister relationship between sandgrouse and pigeons while other studies favored a sister grouping of mesites and sandgrouse instead.
Picodynastornithes is a clade that contains the orders Coraciiformes and Piciformes. This grouping also has current and historical support from molecular and morphological studies.
Columbaves is a clade that contains Columbimorphae and Otidimorphae discovered by genomic analysis by Prum et al. (2015). This conflicts with the Columbea and Otidae hypotheses which Mirandornithes are the sister taxon to Columbimorphae and Cypselomorphae the sister taxon to Otidimorphae, respectively, found by Jarvis et al. (2014). Neither hypothesis supports the two subdivisions of Metaves and Coronoaves as previous studies had found.
Aequorlitornithes is a clade of waterbirds recovered in a comprehensive genomic systematic study using nearly 200 species in 2015. It contains the clades Charadriiformes, Mirandornithes and Phaethoquornithes. Previous studies have found different placement for the clades in the tree.