Comahuesuchidae | |
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Reconstructed skeleton of Comahuesuchus brachybuccalis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauria |
Clade: | Pseudosuchia |
Clade: | Crocodylomorpha |
Clade: | Crocodyliformes |
Clade: | † Notosuchia |
Family: | † Comahuesuchidae Bonaparte, 1991 |
Genera | |
Comahuesuchidae is a family of notosuchian crocodyliforms. Constructed in 1991, it includes the genera Comahuesuchus and Anatosuchus . Among the characteristics that are unique to this family is an external naris that is inset into the tip of the snout. There is also a diastema, or gap between the teeth, at the tip of the upper and lower jaws. Both Anatosuchus and Comahuesuchus have maxillary tooth rows in the upper jaw that extend out and over the dentary tooth rows of the lower jaw. [1]
Several recent phylogenetic analyses of notosuchians have placed Anatosuchus outside of Notosuchia altogether, and therefore outside Comahuesuchidae. If this is true, Comahuesuchus would be the only member of Comahuesuchidae. [2]
Ferugliotherium is a genus of fossil mammals in the family Ferugliotheriidae from the Campanian and/or Maastrichtian period of Argentina. It contains a single species, Ferugliotherium windhauseni, which was first described in 1986. Although originally interpreted on the basis of a single brachydont (low-crowned) molar as a member of Multituberculata, an extinct group of small, rodent-like mammals, it was recognized as related to the hypsodont (high-crowned) Sudamericidae following the discovery of additional material in the early 1990s. After a jaw of the sudamericid Sudamerica was described in 1999, these animals were no longer considered to be multituberculates and a few fossils that were previously considered to be Ferugliotherium were assigned to unspecified multituberculates instead. Since 2005, a relationship between gondwanatheres and multituberculates has again received support. A closely related animal, Trapalcotherium, was described in 2009 on the basis of a single tooth.
Ferugliotheriidae is one of three known families in the order Gondwanatheria, an enigmatic group of extinct mammals. Gondwanatheres have been classified as a group of uncertain affinities or as members of Multituberculata, a major extinct mammalian order. The best-known representative of Ferugliotheriidae is the genus Ferugliotherium from the Late Cretaceous epoch in Argentina. A second genus, Trapalcotherium, is known from a single tooth, a first lower molariform, from a different Late Cretaceous Argentinean locality. Another genus known from a single tooth, Argentodites, was first described as an unrelated multituberculate, but later identified as possibly related to Ferugliotherium. Finally, a single tooth from the Paleogene of Peru, LACM 149371, perhaps a last upper molariform, and a recent specimen from Mexico, may represent related animals.
Doratodon is an extinct genus of Late Cretaceous crocodylomorph that may have been a member of the Sebecosuchia. Doratodon was a relatively small animal with ziphodont teeth, meaning the teeth had flattened sides and serrated edges. Two species of Doratodon are known to science: D. carcharidens from Austria and Hungary, the type species; and D. ibericus from Spain. Teeth similar to those of Doratodon are also known from Italy and Romania, though they cannot be confidently assigned to this genus.
Simosuchus is an extinct genus of notosuchian crocodyliforms from the Late Cretaceous of Madagascar. It is named for its unusually short skull. Fully grown individuals were about 0.75 metres (2.5 ft) in length. The type species is Simosuchus clarki, found from the Maevarano Formation in Mahajanga Province, although one isolated multicuspid tooth of this genus was discovered in Kallamedu Formation of India.
Mahajangasuchus is an extinct genus of crocodyliform which had blunt, conical teeth. The type species, M. insignis, lived during the Late Cretaceous; its fossils have been found in the Maevarano Formation in northern Madagascar. It was a fairly large predator, measuring up to 4 metres (13 ft) long.
Anatosuchus is an extinct genus of notosuchian crocodyliforms discovered in Gadoufaoua, Niger, and described by a team of palaeontologists led by the American Paul Sereno in 2003, in the Journal of Vertebrate Paleontology. Its duck-like snout coincidentally makes it resemble a crocoduck, an imagined hybrid animal with the head of a crocodile and the body of a duck.
Comahuesuchus is an extinct genus of notosuchian crocodylomorphs from the late Cretaceous of Argentina. It was described by palaeontologist José Bonaparte in 1991. The type species is C. brachybuccalis from the Santonian Bajo de la Carpa Formation. In 2023 another species, C. bonapartei from the Sierra Barrosa Formation and Portezuelo Formation, was assigned to the genus.
Razanandrongobe is a genus of carnivorous ziphosuchian crocodyliform from the Middle Jurassic of Madagascar. It contains the type and only species Razanandrongobe sakalavae, named in 2004 by Simone Maganuco and colleagues based on isolated bones found in 2003. The remains, which included a fragment of maxilla and teeth, originated from the Bathonian-aged Sakaraha Formation of Mahajanga, Madagascar. While they clearly belonged to a member of the Archosauria, Maganuco and colleagues refrained from assigning the genus to a specific group because the fragmentary remains resembled lineages among both the theropod dinosaurs and crocodylomorphs.
Notosuchia is a suborder of primarily Gondwanan mesoeucrocodylian crocodylomorphs that lived during the Jurassic and Cretaceous. Some phylogenies recover Sebecosuchia as a clade within Notosuchia, others as a sister group ; if Sebecosuchia is included within Notosuchia its existence is pushed into the Middle Miocene, about 11 million years ago. Fossils have been found from South America, Africa, Asia, and Europe. Notosuchia was a clade of terrestrial crocodilians that evolved a range of feeding behaviours, including herbivory (Chimaerasuchus), omnivory (Simosuchus), and terrestrial hypercarnivory (Baurusuchus). It included many members with highly derived traits unusual for crocodylomorphs, including mammal-like teeth, flexible bands of shield-like body armor similar to those of armadillos (Armadillosuchus), and possibly fleshy cheeks and pig-like snouts (Notosuchus). The suborder was first named in 1971 by Zulma Gasparini and has since undergone many phylogenetic revisions.
Baurusuchidae is a Gondwanan family of mesoeucrocodylians that lived during the Late Cretaceous. It is a group of terrestrial hypercarnivorous crocodilians from South America and possibly Pakistan. Baurusuchidae has been, in accordance with the PhyloCode, officially defined as the least inclusive clade containing Cynodontosuchus rothi, Pissarrachampsa sera, and Baurusuchus pachecoi. Baurusuchids have been placed in the suborder Baurusuchia, and two subfamilies have been proposed: Baurusuchinae and Pissarrachampsinae.
Araripesuchus is a genus of extinct crocodyliform that existed during the Cretaceous period of the late Mesozoic era some 125 to 66 million years ago. Six species of Araripesuchus are currently known. They are generally considered to be notosuchians, characterized by their varied teeth types and distinct skull elements. This genus consists of six species: A. buitreraensis, discovered in Argentina, A. wegeneri, discovered in Cameroon and Niger, A. rattoides, discovered in Niger, A. tsangatsangana, discovered in Madagascar, A. gomesii, discovered in Brazil and another species discovered in Argentina, A. patagonicus. It has been argued that the phylogenetic position of this genus is uncertain, and that taxonomic revision is required.
Candidodon is an extinct genus of notosuchian mesoeucrocodylian. Fossils have been found in the Early Cretaceous Itapecuru Formation in Brazil.
Kaprosuchus is an extinct genus of mahajangasuchid crocodyliform. It is known from a single nearly complete skull collected from the Upper Cretaceous Echkar Formation of Niger. The name means "boar crocodile" from the Greek κάπρος, kapros ("boar") and σοῦχος, soukhos ("crocodile") in reference to its unusually large caniniform teeth which resemble those of a boar. It has been nicknamed "BoarCroc" by Paul Sereno and Hans Larsson, who first described the genus in a monograph published in ZooKeys in 2009 along with other Saharan crocodyliformes such as Anatosuchus and Laganosuchus. The type species is K. saharicus.
Mahajangasuchidae is an extinct family of notosuchian crocodyliforms. It currently contains two genera, Mahajangasuchus and Kaprosuchus, both of which lived during the Late Cretaceous in Gondwana. It is defined as the most inclusive clade containing Mahajangasuchus insignis but not Notosuchus terrestris, Simosuchus clarki, Araripesuchus gomesii, Baurusuchus pachecoi, Peirosaurus torminni, Goniopholis crassidens, Pholidosaurus schaumbergensis, or Crocodylus niloticus. Phylogenetically, Mahajangasuchidae is placed just outside pholidosaurids and more derived neosuchians.
Yacarerani is an extinct genus of Late Cretaceous notosuchian crocodylomorph. Fossils have been found in 2002 in the Cajones Formation in Amboró National Park of central Bolivia and are Turonian-Santonian in age. The genus was described in 2009 in the Journal of Vertebrate Paleontology. The material represents two individuals and was found in association with eggs that are thought to have been part of a nest. Yacarerani was a small crocodylomorph at around 80 centimetres (31 in) in length. It may have lived in small groups, creating burrows to lay eggs in.
Sebecus is an extinct genus of sebecid crocodylomorph from Eocene of South America. Like other sebecosuchians, it was entirely terrestrial and carnivorous. The genus is currently represented by two species, the type S. icaeorhinus and S. ayrampu. Several other species have been referred to Sebecus, but were later reclassified as their own genera.
Aetodactylus is a genus of targaryendraconian pterosaur. It is known from a lower jaw discovered in Upper Cretaceous rocks of northeastern Texas, United States.
Candidodontidae is a family of notosuchian crocodyliforms. It was originally used in 2002 as a name for a clade that includes the genera Araripesuchus, Candidodon, and Malawisuchus. Later in 2004 the family was formally defined as a node-based taxon including Candidodon itapecuruense and Mariliasuchus amarali. A 2009 study redefined Candidodontidae as a stem-based taxon which included Candidodon, Malawisuchus, and possibly Mariliasuchus.
Gwawinapterus beardi is a species of saurodontid ichthyodectiform fish from the Late Cretaceous period of British Columbia, Canada. While initially described as a very late-surviving member of the pterosaur family Istiodactylidae, further examination has cast doubt on the identification of the specimen as a pterosaur, and research published in 2012 identified the remains as having come from a saurodontid fish.
Campinasuchus is an extinct genus of baurusuchid mesoeucrocodylian from Minas Gerais State of Brazil.