Condylodon | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Family: | Formicidae |
Subfamily: | incertae sedis |
Genus: | Condylodon Lund, 1831 |
Species: | C. audouini |
Binomial name | |
Condylodon audouini Lund, 1831 [1] | |
Condylodon is a genus of ant with an uncertain placement ( incertae sedis ) in the family Formicidae. [2] It contains the single species Condylodon audouini, first described from a single specimen by Lund (1831) in a paper on Brazilian ants. [3]
The type material is presumed to be lost and the taxonomy of species remains uncertain. It has variously been speculated to belong to different families and subfamilies, including Mutillidae, a family of wasps with females resembling large ants, by Swainson & Shuckard (1840). Dalla Torre (1893) relegated the genus to a junior synonym of Pseudomyrma (now Pseudomyrmex , subfamily Pseudomyrmecinae). Emery (1921) dubiously placed it in Ponerinae. Ward (1990) excluded the taxon from Pseudomyrmecinae, and agreed with Emery's placement in Ponerinae as the most likely candidate, and listed Ectatomma as a possible identity of the genus. Bolton (1994) placed it as incertae sedis in Ponerinae, and most recently, as incertae sedis in Formicidae (Bolton, 2003). [2] [3]
Ponerinae, the ponerine ants, is a subfamily of ants in the Poneromorph subfamilies group, with about 1,600 species in 47 extant genera, including Dinoponera gigantea - one of the world's largest species of ant. Mated workers have replaced the queen as the functional egg-layers in several species of ponerine ants. In such queenless species, the reproductive status of workers can only be determined through ovarian dissections.
Myrmicinae is a subfamily of ants, with about 140 extant genera; their distribution is cosmopolitan. The pupae lack cocoons. Some species retain a functional sting. The petioles of Myrmicinae consist of two nodes. The nests are permanent and in soil, rotting wood, under stones, or in trees.
Myrmeciinae is a subfamily of the Formicidae, ants once found worldwide but now restricted to Australia and New Caledonia. This subfamily is one of several ant subfamilies which possess gamergates, female worker ants which are able to mate and reproduce, thus sustaining the colony after the loss of the queen. The Myrmeciinae subfamily was formerly composed of only one genus, Myrmecia, but the subfamily was redescribed by Ward & Brady in 2003 to include two tribes and four genera. An additional three genera, one form genus, and 9 species were described in 2006 from the Early Eocene of Denmark, Canada, and Washington. Subsequently an additional fossil genus was moved from the family Rhopalosomatidae in 2018, and a new genus was described in 2021.
Dolichoderinae is a subfamily of ants, which includes species such as the Argentine ant, the erratic ant, the odorous house ant, and the cone ant. The subfamily presents a great diversity of species throughout the world, distributed in different biogeographic realms, from the Palearctic, Nearctic, Afrotropical region and Malaysia, to the Middle East, Australian, and Neotropical regions.
Pseudomyrmecinae is a small subfamily of ants containing only three genera of slender, large-eyed arboreal ants, predominantly tropical or subtropical in distribution. In the course of adapting to arboreal conditions, the pseudomyrmecines diversified and came to occupy and retain a much wider geographic range.
Pachycondyla is a ponerine genus of ants found in the Neotropics.
Aneuretinae is a subfamily of ants consisting of a single extant species, Aneuretus simoni, and 9 fossil species. Earlier, the phylogenetic position of A. simoni was thought to be intermediate between primitive and advanced subfamilies of ants, but recent studies have shown it is the nearest living relative of subfamily Dolichoderinae.
Ypresiomyrma is an extinct genus of ants in the subfamily Myrmeciinae that was described in 2006. There are four species described; one species is from the Isle of Fur in Denmark, two are from the McAbee Fossil Beds in British Columbia, Canada, and the fourth from the Bol’shaya Svetlovodnaya fossil site in Russia. The queens of this genus are large, the mandibles are elongated and the eyes are well developed; a stinger is also present. The behaviour of these ants would have been similar to that of extant Myrmeciinae ants, such as solitary foraging for arthropod prey and never leaving pheromone trails. The alates were poor flyers due to their size, and birds and animals most likely preyed on these ants. Ypresiomyrma is not assigned to any tribe, and is instead generally regarded as incertae sedis within Myrmeciinae. However, some authors believe Ypresiomyrma should be assigned as incertae sedis within Formicidae.
Macabeemyrma is an extinct genus of bulldog ants in the subfamily Myrmeciinae containing the single species Macabeemyrma ovata, described in 2006 from Ypresian stage deposits of British Columbia, Canada. Only a single specimen is known; a holotype queen found preserved as a compression fossil. The specimen had no wings and small portions of its legs and eyes were faintly preserved. It was a large ant, reaching 25 millimetres (0.98 in) in length. This ants' behaviour would have been similar to that of extant Myrmeciinae ants, such as foraging singly in search for arthropod prey and nesting in soil or in trees. Macabeemyrma shows similarities to extinct ants in the genus Ypresiomyrma, and to the living Nothomyrmecia macrops, but has not been conclusively assigned to any tribe, instead generally regarded as incertae sedis within Myrmeciinae. However, the sole specimen lacks definitive traits, and its classification in Myrmeciinae, and even its identity as an ant, has been challenged.
Myrmeciites is an extinct form genus of bulldog ants in the subfamily Myrmeciinae of the family Formicidae, which contains three described species and two fossils not placed beyond the genus level. Described in 2006 from Ypresian stage deposits, all three of the described species and one unplaced fossil are from British Columbia, Canada, while the second unplaced fossil is from Washington State, USA. These ants were large, with the largest specimens collected reaching 3 centimetres (1.2 in). The behaviour of these ants would have been similar to extant Myrmeciinae ants, such as solitary foraging, nesting either in the soil or trees, and leaving no pheromone trail to food sources. Due to the poor preservation of these ants, their phylogenetic position among Myrmeciinae is unclear, and no type species has been designated. These ants are classified as incertae sedis in Myrmeciinae, but some writers have classified it as incertae sedis within the insect order Hymenoptera. This reclassification, however, has not been accepted; instead, Myrmeciites remains in Myrmeciinae.
Sphecomyrminae is an extinct subfamily of ants in family Formicidae known from a series of Cretaceous fossils found in North America, Europe, and Asia. Sphecomyrminae contains eight genera, divided into two tribes Sphecomyrmini and Zigrasimeciini. The tribe Sphecomyrmini contains the six genera Armania, Cretomyrma, Gerontoformica, Orapia, Pseudarmania and Sphecomyrma; while Zigrasimeciini contains Boltonimecia and Zigrasimecia. A number of taxa have been removed from the subfamily and placed either in other subfamilies or are now treated as incertae sedis in Formicidae.
Brownimecia is an extinct genus of ants, the only genus in the tribe Brownimeciini and subfamily Brownimeciinae of the Formicidae. Fossils of the identified species, Brownimecia clavata and Brownimecia inconspicua, are known from the Late Cretaceous of North America. The genus is one of several ants described from Late Cretaceous ambers of New Jersey. Brownimecia was initially placed in the subfamily Ponerinae, until it was transferred to its own subfamily in 2003; it can be distinguished from other ants due to its unusual sickle-like mandibles and other morphological features that makes this ant unique among the Formicidae. B. clavata is also small, measuring 3.43 millimetres (0.135 in), and a stinger is present in almost all of the specimens collected. The morphology of the mandibles suggest a high level of feeding specialization.
Meranoplus parviumgulatus is a species of ant in the genus Meranoplus. It is known from New Guinea and Papua New Guinea.
Burmomyrma is an extinct genus of aculeate hymenopteran, suggested to be an ant. The genus contains a single described species, Burmomyrma rossi. Burmomyrma is known from a single Middle Cretaceous fossil which was found in Asia.
Archimyrmex is an extinct genus of ant in the formicid subfamily Myrmeciinae, described by palaeoentomologist Theodore Cockerell in 1923. The genus contains four described species, Archimyrmex rostratus, Archimyrmex piatnitzkyi, Archimyrmex smekali and Archimyrmex wedmannae. Archimyrmex is known from a group of Middle Eocene fossils which were found in North America, South America, and Europe. The genus was initially placed in the subfamily Ponerinae, but it was later placed in Myrmeciinae; it is now believed to be the ancestor of the extant primitive genus Myrmecia from Australia. Despite this, Archimyrmex is not a member to any tribe and is regarded as incertae sedis within Myrmeciinae. However, some authors believe Archimyrmex should be assigned as incertae sedis within Formicidae. These ants can be characterised by their large mandibles and body length, ranging from 13.2 to 30 mm. They also have long, thin legs and an elongated mesosoma (thorax) and petiole.
Armaniinae is subfamily of extinct ant-like hymenopterans known from a series of Cretaceous fossils found in Asia and Africa. It is usually treated as one of the stem-group subfamilies in family Formicidae, although some myrmecologists treat it as a distinct family. A 2007 study analysing petiole and antenna morphology led to the proposal that at least some of the armaniid genera be placed in Sphecomyrminae, although others are unconvinced by the arguments and retain Armaniinae. The subfamily contains seven genera with fourteen described species.
Noonilla is a genus of ants of uncertain placement in the family Formicidae. It contains the single species Noonilla copiosa, first described by Petersen in 1968 based on male specimen from the Philippines. Noonilla was initially placed in the subfamily Leptanillinae, but was later removed from the subfamily when Ogata, Terayama & Masuko (1995) reviewed the genus, leaving the genus incertae sedis in the family.
Pachycondyla succinea is an extinct species of ant in the formicid subfamily Ponerinae described from fossils found in Europe. P. petrosa is one of three middle Eocene Pachycondyla species found in Baltic amber.