Epileolis Temporal range: | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Squamata |
Suborder: | Iguania |
Family: | Agamidae |
Genus: | † Epileolis Alifanov, 2020 |
Species: | †E. reshetovi |
Binomial name | |
†Epileolis reshetovi Alifanov, 2020 | |
Epileolis is an extinct genus of agamid lizard that inhabited Mongolia during the Late Palaeocene. It contains a single species, Epileolis reshetovi. [1]
The Phrynosomatidae are a diverse family of lizards, sometimes classified as a subfamily (Phrynosomatinae), found from Panama to the extreme south of Canada. Many members of the group are adapted to life in hot, sandy deserts, although the spiny lizards prefer rocky deserts or even relatively moist forest edges, and the short-horned lizard lives in prairie or sagebrush environments. The group includes both egg-laying and viviparous species, with the latter being more common in species living at high elevations.
The Varanidae are a family of lizards in the superfamily Varanoidea and order Anguimorpha. The family, a group of carnivorous and frugivorous lizards, includes the living genus Varanus and a number of extinct genera more closely related to Varanus than to the earless monitor lizard (Lanthanotus). Varanus includes the Komodo dragon, crocodile monitor, savannah monitor, the goannas of Australia and Southeast Asia, and various other species with a similarly distinctive appearance. Their closest living relatives are the earless monitor lizard and Chinese crocodile lizard. The oldest members of the family are known from the Late Cretaceous of Mongolia.
Estesia is an extinct genus of Late Cretaceous anguimorph lizard found in the Gobi Desert in Mongolia.
Gobiderma is an extinct genus of Late Cretaceous lizard whose fossils are known from the Gobi Desert in southern Mongolia. It was first discovered as a result of a joint Polish-Mongolian Paleontological Expedition, and formally named in 1984. In life, it probably resembled lizards of the genus Heloderma to a large degree, though its skull was more elongated than lizards of that genus.
Dorsetisaurus is a genus of extinct lizard, known from the Late Jurassic of North America, and the Late Jurassic-earliest Cretaceous of Europe. The genus was first reported from the Early Cretaceous (Berriasian) Lulworth Formation of the Purbeck Group of Durlston Bay, in Dorset. It has also been reported from the Late Jurassic aged Alcobaça Formation of Portugal, the Aptian-Albian Dzunbain Formation of Mongolia, and the Morrison Formation of Western North America present in stratigraphic zones 2, 4, and 5. It is considered the oldest widely accepted member of Anguimorpha. based on the presence of 11 shared synapomorphies.
Isodontosaurus is an extinct genus of iguanian lizard from the Late Cretaceous of Mongolia and China. The type species is Isodontosaurus gracilis. Isodontosaurus is part of an extinct group of Late Cretaceous iguanians called Gobiguania, which is currently thought to be endemic to Mongolia.
Ovoo gurvel is an extinct varanid lizard from the Late Cretaceous of Mongolia. It is one of the smallest and earliest monitor lizards. It was described in 2008. Ovoo possesses a pair of small bones in its skull that are not seen in any other lizard.
Aiolosaurus is an extinct genus of monitor lizard from the Late Cretaceous of Mongolia. The type and only species, Aiolosaurus oriens, was named in 2000 from Ukhaa Tolgod, a rich fossil site in the Campanian-age Djadochta Formation.
Cherminotus is an extinct genus of monitor lizard from the Late Cretaceous of Mongolia. The type and only species, Cherminotus longifrons, was named in 1984.
Gobekko is an extinct genus of gecko or gecko-like lizard from the Late Cretaceous Djadokhta Formation of the Gobi Desert in Mongolia. Gobekko is either a basal member of Gekkota, the group that includes geckos and the legless pygopodid lizards, or a stem-gekkotan outside Gekkota but within the larger group Gekkonomorpha. It is the fourth oldest known member of Gekkonomorpha after Hoburogekko, a gecko from the Early Cretaceous of Mongolia, AMNH FR21444, an unnamed specimen also from the Early Cretaceous of Mongolia, and Cretaceogekko, a gecko preserved in amber from the Early Cretaceous of Burma.
Desertiguana is an extinct genus of lizard in the family Phrynosomatidae. It is a monotypic genus represented by the type species Desertiguana gobiensis from the Late Cretaceous Barun Goyot Formation of Mongolia. Desertiguana gobiensis is known from a single left lower jaw.
Gobiguania is an extinct clade of iguanian lizards from the Late Cretaceous. All known gobiguanians are endemic to the Gobi Desert of Mongolia. Gobiguania was given a phylogenetic definition by Jack Conrad and Mark Norell in 2007 as all taxa more closely related to Anchaurosaurus gilmorei than to Iguana iguana, Crotaphytus collaris, or Agama agama. According to Conrad and Norell's phylogenetic analysis, Gobiguania includes Anchaurosaurus as well as several other Late Cretaceous lizards such as Ctenomastax, Temujinia, Saichangurvel, and Zapsosaurus. A phylogenetic analysis published in 2012 indicated that three other lizard genera — Igua, Isodontosaurus, and Polrussia, all from Mongolia and all from the Late Cretaceous — are also gobiguanians. Below is a cladogram from the analysis:
Priscagamidae is an extinct family of iguanian lizards known from the Late Cretaceous of Mongolia and China and the Eocene of India, spanning a range from 83.6 to 48.6 million years ago. Probably the earliest priscagamids on indeterminate genera were found in Aptian-Albian sediments in "Hobur", Mongolia. It includes the genera Heterodontagama, Mimeosaurus, Phrynosomimus, Priscagama, and possibly Pleurodontagama. The first fossils of priscagamids were found in the Djadochta and Khermeen Tsav formations of Mongolia. More recently they have been found in the Cambay Formation in India, leading to the naming of Heterodontagama in 2013. Priscagamidae was originally described as a subfamily of Agamidae called Priscagaminae in 1984, but it was reclassified as a distinct family in 1989. Most phylogenetic analyses still find a close relationship between Priscagamidae and Agamidae, although a 2015 study found it to be basal to all other iguanian clades, warranting its removal from Iguania and placement in a larger clade called Iguanomorpha.
Priscagama is an extinct genus of iguanian lizard from the Late Cretaceous of Mongolia and China. It belongs to an extinct family of iguanians called Priscagamidae. Several incomplete skulls have been found in the Barun Goyot and Djadochta formations, and were originally referred to the genus Mimeosaurus; the type species Priscagama gobiensis was named in 1984 when it was recognized that these skulls belonged to a distinct species. Priscagama differs from most other priscagamids in having a more elongate, lightly built skull. It is very similar in appearance to another priscagamid called Pleurodontagama, as the two can only be distinguished by the shape of their teeth.
Phrynosomimus is an extinct genus of iguanian lizard from the Late Cretaceous of Mongolia belonging to the extinct family Priscagamidae. The type species Phrynosomimus asper was named in 1996. Fossils have been found in the Barun Goyot and Djadochta formations and include several complete skulls. Phrynosomimus has a short, triangular skull with bony spikes projecting from the back, stemming from the squamosal and parietal bones. These spikes give it a similar appearance to the modern horned lizard Phrynosoma and inspire its name, which means "Phrynosoma mimic." Like other priscagamids it has an acrodont dentition, meaning that its teeth grow from the margins of the jaws rather than their inner surfaces, as is the case for the pleurodont dentitions of most lizards.
Anchaurosaurus is an extinct genus of iguanian lizard from the Late Cretaceous of Inner Mongolia, China. It belongs to an extinct clade of iguanians called Gobiguania that was endemic to the Gobi Desert during the Late Cretaceous. The type species, Anchaurosaurus gilmorei, was named in 1995 on the basis of a well-preserved skull and incomplete skeleton from the Djadochta Formation. Compared to other iguanians, Anchaurosaurus has a relatively elongated skull, large eye sockets, and higher tooth crowns. Phylogenetic analysis indicates that among gobiguanians, Anchaurosaurus is most closely related to Zapsosaurus from Mongolia. Below is a cladogram from Daza et al. (2012) showing the phylogenetic relationships of Anchaurosaurus:
Xihaina is an extinct genus of iguanian lizard from the Late Cretaceous of Inner Mongolia, China. The type species Xihaina aquilonia was named in 1995 from the Djadochta Formation and is known from a partial skeleton that preserves parts of the skull, most of the vertebral column, the pelvis, and the right hind limb. The incomplete nature of this specimen makes the classification of Xihaina difficult; it has never been incorporated into a phylogenetic analysis, but it shares similarities with a group of Late Cretaceous Mongolian lizards called Gobiguania, particularly the gobiguanian genera Anchaurosaurus and Polrussia. The fact that many skeletal elements are missing yet the rest of the skeleton is articulated suggests the individual may have been partially eaten by a predator or scavenger and then rapidly buried soon after.
Polrussia is an extinct genus of iguanian lizards dating to the Late Cretaceous epoch, found in what is now Mongolia. It belongs to a group of extinct iguanians called Gobiguania that was endemic to the Gobi Desert during the Late Cretaceous. The type species Polrussia mongoliensis was named in 1991 on the basis of a skull found in the Barun Goyot Formation. The genus name refers to the Polish and Russian paleontologists who worked together to find and describe the material. Polrussia has a short skull, slightly pointed and flattened snout, and large eye sockets. The teeth each have one cusp, as opposed to the multiple cusps seen in some other gobiguanians. The skull is only 1.2 centimetres (0.47 in) long, making Polrussia one of the smallest gobiguanians.
Igua is an extinct genus of iguanian lizards belonging to a group called Gobiguania that was endemic to the Gobi Desert during the Late Cretaceous. The type species Igua minuta was named in 1991 on the basis of a skull from the Barun Goyot Formation in Mongolia. The skull itself is very small, only 14 millimetres (0.55 in) long, and may have belonged to a juvenile given that it possesses a large fontanelle and that many of the bones are unfused. The snout-vent length of the individual is estimated to have been 55 to 65 millimetres. Igua differs from related gobiguanians like Polrussia in having a more rounded skull. It is similar in appearance to the living genera Liolaemus and Tropidurus. The teeth are tricuspid and pleurodont, meaning they are attached to inner surfaces of the jaws.
Ondogurvel is a genus of alvarezsaurid dinosaur from the Late Cretaceous (Campanian) Barun Goyot Formation in southern Mongolia. The type and only species is O. alifanovi, known from a partial skeleton consisting of fragments of two last dorsal vertebrae, three anterior sacral vertebrae, right ilium, left and right pubis and ischium, articulated right tibia, fibula, metatarsals II and IV, and phalanges IV-1 and IV-2, right carpometacarpus, left and right manual phalanx II-1, right femur, left pedal phalanx II-1, and fragments of unidentified phalanges.