Escuminac Formation | |
---|---|
Stratigraphic range: Devonian (Middle Frasnian) | |
Type | Geological formation |
Unit of | Miguasha Group [1] |
Underlies | Bonaventure Formation [1] |
Overlies | Fleurant Formation [1] |
Location | |
Region | Quebec |
Country | Canada |
The Escuminac Formation is a geologic formation in Quebec. It preserves fossils dating back to the Frasnian, in the Devonian period. [2] [3]
Miguasha National Park is located within this formation along the estuary of the Restigouche River on the south coast of the Gaspé Peninsula. The fossil biota from the Park is thus referred to as the Miguasha biota. [1] The main Miguasha exposures were named the 'René Bureau Cliffs' after the geologist and paleontologist. [1] This formation is sufficiently rich that some material could be sacrificed for microanatomical and histological study through thin sectioning; this was done, among others, for the actinopterygian Cheirolepis , [4] the actinistian Miguashaia [5] and the tetrapodomorph Euthenopteron . [6]
The formation's depositional environment has been variously considered as lacustrine, estuarine, coastal marine or marine, though evidence from the fossil assemblage, stratigraphic and sedimentological setting, and geochemistry of the sedimentary rocks and bones suggests an estuarine interpretation is most fitting. [1]
Acanthodians | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Diplacanthus | D. ellsi | [1] | A diplacanthiform. | ||
D. horridus | [1] | A diplacanthiform. | |||
Homalacanthus | H. concinnus | [1] | |||
Triazeugacanthus | T. affinis | Miguasha, Quebec. [1] [7] | Thousands of specimens representing multiple ontogenetic stages. [7] | ||
Actinopterygians | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Cheirolepis | C. canadensis | [1] | |||
Jawless fish | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Endeiolepis | E. aneri | [1] | |||
Escuminaspis | E. laticeps | [1] | |||
Euphanerops | E. longaevus | Miguasha. [1] [8] | Over 3,500 specimens. [8] | A euphaneropid. | |
Levesquaspis | L. patteni | [1] | |||
Placoderms | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Bothriolepis | B. canadensis | [1] | An antiarch. | ||
Plourdosteus | P. canadensis | [1] | An arthrodire. | ||
Sarcopterygians | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Callistiopterus | C. clappi | [1] | |||
Elpistostege | E. watsoni | [1] | A stegocephalian. | ||
Eusthenopteron | E. foordi | [1] | A tristichopterid. | ||
Fleurantia | F. denticulata | [1] | A lungfish. | ||
Holoptychiidae | Holoptychiidae nov. sp. | [1] | A porolepiform. | ||
Holoptychius | H. jarviki | [1] | A porolepiform. | ||
Miguashaia | M. bureaui | Miguasha, Quebec. [1] [9] | A coelacanth. | ||
Quebecius | Q. quebecensis | [1] | A porolepiform. | ||
Scaumenacia | S. curta | [1] | A lungfish. | ||
Arthropods | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Asmusia | A. membranacea | [1] | A clam shrimp. | ||
Pterygotus | P. sp. | [10] | An eurypterid. | ||
Petaloscorpio | P. bureaui | [1] | A scorpion. | ||
Zanclodesmus | Z. willetti | [1] [11] | A millipede. | ||
Ctenophores | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Daihuoides | D. jakobvintheri | Miguasha, Quebec. [12] | MHNM 24–01. [12] | ||
Plants | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Archaeopteris | A. halliana | [1] | |||
Barynophyton | [1] | ||||
Flabellofolium | [1] | ||||
Protobarynophyton | [1] | ||||
Spermasporites | [1] | ||||
Sarcopterygii — sometimes considered synonymous with Crossopterygii — is a clade of bony fish commonly referred to as lobe-finned fish. They are characterised by prominent muscular limb buds (lobes) within their fins, which are supported by articulated appendicular skeletons. This is in contrast to the other clade of bony fish, the Actinopterygii, which have only skin-covered bony spines supporting the fins.
Tooth enamel is one of the four major tissues that make up the tooth in humans and many animals, including some species of fish. It makes up the normally visible part of the tooth, covering the crown. The other major tissues are dentin, cementum, and dental pulp. It is a very hard, white to off-white, highly mineralised substance that acts as a barrier to protect the tooth but can become susceptible to degradation, especially by acids from food and drink. In rare circumstances enamel fails to form, leaving the underlying dentin exposed on the surface.
Dentin or dentine is a calcified tissue of the body and, along with enamel, cementum, and pulp, is one of the four major components of teeth. It is usually covered by enamel on the crown and cementum on the root and surrounds the entire pulp. By volume, 45% of dentin consists of the mineral hydroxyapatite, 33% is organic material, and 22% is water. Yellow in appearance, it greatly affects the color of a tooth due to the translucency of enamel. Dentin, which is less mineralized and less brittle than enamel, is necessary for the support of enamel. Dentin rates approximately 3 on the Mohs scale of mineral hardness. There are two main characteristics which distinguish dentin from enamel: firstly, dentin forms throughout life; secondly, dentin is sensitive and can become hypersensitive to changes in temperature due to the sensory function of odontoblasts, especially when enamel recedes and dentin channels become exposed.
Acanthodii or acanthodians is an extinct class of gnathostomes. They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans and chondrichthyans. In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians.
Pterygotus is a genus of giant predatory eurypterid, a group of extinct aquatic arthropods. Fossils of Pterygotus have been discovered in deposits ranging in age from Middle Silurian to Late Devonian, and have been referred to several different species. Fossils have been recovered from four continents; Australia, Europe, North America and South America, which indicates that Pterygotus might have had a nearly cosmopolitan (worldwide) distribution. The type species, P. anglicus, was described by Swiss naturalist Louis Agassiz in 1839, who gave it the name Pterygotus, meaning "winged one". Agassiz mistakenly believed the remains were of a giant fish; he would only realize the mistake five years later in 1844.
Eusthenopteron is a genus of prehistoric sarcopterygian fishes known from several species that lived during the Late Devonian period, about 385 million years ago. It has attained an iconic status from its close relationship to tetrapods. Early depictions of animals of this genus show them emerging onto land, but paleontologists now widely agree that eusthenopteron species were strictly aquatic animals.
Miguasha Provincial Park is a protected area near Carleton-sur-Mer on the Gaspé Peninsula of Quebec in Canada. Created in 1985 by the Government of Quebec, Miguasha was designated a World Heritage Site in 1999 in recognition of its wealth of fossils, which display a crucial time during the evolution of life on Earth. Other names for this site are the Miguasha Fossil Site, the Bay of Escuminac Fossil Site, the Upper Devonian Escuminac Formation, and the Hugh-Miller Cliffs. It is also sometimes referred to on fossil specimens as 'Scaumenac Bay' or 'Scaumenac Bay P.Q.'
Bothriolepis was a widespread, abundant and diverse genus of antiarch placoderms that lived during the Middle to Late Devonian period of the Paleozoic Era. Historically, Bothriolepis resided in an array of paleo-environments spread across every paleocontinent, including near shore marine and freshwater settings. Most species of Bothriolepis were characterized as relatively small, benthic, freshwater detritivores, averaging around 30 centimetres (12 in) in length. However, the largest species, B. rex, had an estimated bodylength of 170 centimetres (67 in). Although expansive with over 60 species found worldwide, comparatively Bothriolepis is not unusually more diverse than most modern bottom dwelling species around today.
Tetrapodomorpha is a clade of vertebrates consisting of tetrapods and their closest sarcopterygian relatives that are more closely related to living tetrapods than to living lungfish. Advanced forms transitional between fish and the early labyrinthodonts, such as Tiktaalik, have been referred to as "fishapods" by their discoverers, being half-fish, half-tetrapods, in appearance and limb morphology. The Tetrapodomorpha contains the crown group tetrapods and several groups of early stem tetrapods, which includes several groups of related lobe-finned fishes, collectively known as the osteolepiforms. The Tetrapodomorpha minus the crown group Tetrapoda are the stem Tetrapoda, a paraphyletic unit encompassing the fish to tetrapod transition.
Cheirolepis is an extinct genus of ray-finned fish that lived in the Devonian period of Europe and North America. It is the only genus yet known within the family Cheirolepidae and the order Cheirolepiformes. It was among the most basal of the Devonian actinopterygians and is considered the first to possess the "standard" dermal cranial bones seen in later actinopterygians.
Tristichopterids (Tristichopteridae) were a diverse and successful group of fish-like tetrapodomorphs living throughout the Middle and Late Devonian. They first appeared in the Eifelian stage of the Middle Devonian. Within the group sizes ranged from a few tens of centimeters (Tristichopterus) to several meters.
The Fram Formation is an Upper Devonian (Frasnian) sequence of rock strata on Ellesmere Island that came into prominence in 2006 with the discovery in its rocks of examples of the transitional fossil, Tiktaalik, a sarcopterygian or lobe-finned fish showing many tetrapod characteristics. Fossils of Laccognathus embryi, a porolepiform lobe-finned fish, and Qikiqtania, a close relative of Tiktaalik, were also found in the formation. The Fram Formation is a Middle to Upper Devonian clastic wedge forming an extensive continental facies consisting of sediments derived from deposits laid down in braided stream systems that formed some 375 million years ago, at a time when the North American craton ("Laurentia") was straddling the equator.
Elpistostege is an extinct genus of finned tetrapodomorphs that lived during the Frasnian age of the Late Devonian epoch. Its only known species, E. watsoni, was first described in 1938 by the British palaeontologist Thomas Stanley Westoll, based on a single partial skull roof discovered at the Escuminac Formation in Quebec, Canada.
Guiyu oneiros is one of the earliest articulated bony fish discovered. Fossils of Guiyu have been found in what is now Qujing, Yunnan, China, in late Silurian marine strata, about 425 million years old.
Cosmine is a spongy, bony material that makes up the dentine-like layers in the scales of the lobe-finned fishes of the class Sarcopterygii. Fish scales that include layers of cosmine are known as cosmoid scales.
Zanclodesmus willetti is an extinct species of archipolypodan millipede that lived in the Late Devonian period of North America, approximately 380 million years ago. It was described in 2005 based on a fossil discovered in the Escuminac Formation of Quebec, Canada two years prior. It was approximately 44 mm (1.7 in) long and 10 mm wide with 27 body segments, and had kidney shaped patches of ocelli. Each trunk segment had long, sickle-shaped extensions (paranota) projecting laterally, and was decorated on the dorsal surface with low rectangular bosses ("bumps") bordered by crescent-shaped bosses. The genus name Zanclodesmus derives from Greek Zanklon, meaning "sickle", in reference to the long, curved paranota, and desmus, a common root word in millipedes. The species name willietti honors Miguasha National Park warden Jason Willett, who discovered the fossil.
This list of fossil fishes described in 2017 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes and other fishes of every kind that are scheduled to be described during the year 2017, as well as other significant discoveries and events related to paleontology of fishes that are scheduled to occur in the year 2017. The list only includes taxa at the level of genus or species.
The Intertemporal bone is a paired cranial bone present in certain sarcopterygians and extinct amphibian-grade tetrapods. It lies in the rear part of the skull, behind the eyes.
Triazeugacanthus is an extinct genus of spiny shark from the Devonian of Canada. It contains a single species, Triazeugacanthus affinis. With total length up to 5.272 centimetres (2.076 in), it is known from multiple ontogenetic stages.