Centipedes are predatory arthropods belonging to the class Chilopoda of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (metameric) creatures with one pair of legs per body segment. All centipedes are venomous and can inflict painful stings, injecting their venom through pincer-like appendages known as forcipules or toxicognaths, which are actually modified legs instead of fangs. Despite the name, no centipede has exactly 100 pairs of legs; number of legs ranges from 15 pairs to 191 pairs, always an odd number.
Geophilus flavus is a terrestrial, soil-dwelling, species of centipede in the Geophilidae family. G. flavus occurs in a range of habitats across central Europe, North America, Australia and other tropical regions. Geophilomorph centipedes, like centipedes generally, are primary predators, hunting predominantly in underground soil burrows or above ground leaf litter. Their consumption behaviours are influenced by environment and seasonal factors. Given their lack of economic value and marginal medical significance, G.flavus remains largely understudied in mainstream research. Some recent studies have detailed the evolutionary development of G.flavus and Geophilidae generally, illustrating developed predatory features like forcipule venom glands.
The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae.
Geophilus is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with Brachygeophilus. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, anterior porefields, complete or nearly complete coxo-pleural sutures at the prosternum, and incomplete chitin-lines. Centipedes in this genus range from 1 cm to 8 cm in length. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as Geophilus electricus.
Geophilus electricus is a species of soil centipede in the family Geophilidae found across temperate Europe, especially Britain and Ireland. Originally discovered and named Scolopendra electrica in 1758, it was later moved to its own genus, Geophilus. As the first species described in this genus, it is considered by some to be the type species. It is yellowish-orange in color and can grow up to 45 millimeters, with 55 to 75 leg pairs, a distinct carpophagus fossae on the anterior sternites, and a distinctive arrangement of the coxal pores of the last leg pair. Like many geophilomorphans, G. electricus has been known to glow in the dark, most likely as a defensive maneuver.
Geophilus carpophagus is a species of soil centipede in the family Geophilidae, widely considered to be a type species of the genus Geophilus. It grows up to 60 millimeters in length, with an orange/tan body bearing a distinctive purplish marbled pattern. Males of this species have 51 to 55 pairs of legs; females have 53 to 57.
Geophilus easoni is a species of soil centipede in the family Geophilidae found throughout Britain and Ireland, though its range extends through western France to at least the foothills of the Pyrenees. Until 2001, it was considered synonymous with G. carpophagus, and most pre-2001 records probably refer to G. easoni. It's typically shorter than G. carpophagus, with fewer leg pairs, uniform tan/chestnut coloring, and a greater size and number of coxal pores, as well as a darkly pigmented mid-piece of the labrum which bears blunt teeth. Specimens in north-western Iberia typically have a wider range of leg pairs. It nests on the ground surface under stones and dead wood.
Geophilus proximus is a species of soil centipede in the family Geophilidae found in the northern part of the Palearctic and widespread across the entire Baltic basin, though it reaches as far as the Arctic Circle and has been introduced through human agency to northern, central, and eastern parts of Kazakhstan. It was recorded once with certainty in Britain from Unst in the Shetland Islands; distribution in the rest of Europe is difficult to assess because of frequent misidentifications of the species. Populations from northern Europe are mostly parthenogenetic.
Geophilus impressus is a species of soil centipede in the family Geophilidae found all over Europe, and has also been recorded in North Africa. It lives frequently in endogean habitats; in Sardinia it's found mostly in Quercus ilex woods, but also in Mediterranean shrub, open habitats, and maquis. It lives anywhere from sea level to 1700 meters above it, sometimes in caves.
Geophilus gavoyi is a species of soil centipede in the family Geophilidae. It is closely related to both Geophilus aetnensis and its junior synonym G. evisensis; it's considered by some to be a synonym of G. evisensis. It's found under stones in northern Asia and western Europe, especially France and England. It grows to between 23 and 35 millimeters in length and has between 39 and 43 leg pairs in males and 41–45 in females. In the since debunked elongata subspecies it was said to vary from between 49 and 55 leg pairs in males and between 51 and 57 in females. As described in 1964, it has well-developed sternal grooves, longer legs than G. aetnensis, and normal claws in the anal legs.
Geophilus aetnensis is a species of soil centipede in the family Geophilidae found in Europe and northern Asia, excluding China. As described by Verhoeff in 1928, it grows up to 28 millimeters and has 53 leg pairs, 4 sensory setae each on the 2nd-4th front sternites, slightly notched maxillae, and very faint sternal pits on the thorax as well as poorly developed sternal grooves.
Geophilus gracilis is a species of soil centipede in the family Geophilidae found most commonly in Britain and Ireland, though specimens have also been recorded in Chile, France, Greece, and Algeria. It lives under mud and stones along the coast near or below the high tide mark, grows up to 30 millimeters in length, and is bright yellow in color with a darker reddish head. Males of this species have 51 to 57 pairs of legs; females have 51 to 61 leg pairs. This species is often confused with G. flavus and, in coastal locations, G. osquidatum.
Geophilus virginiensis is a species of soil centipede in the family Geophilidae found in Virginia. It grows up to 50 millimeters in length, has 49–57 leg pairs, and is tawny yellow in color.
Geophilus vittatus, also known as the diamondback soil centipede, is a species of soil centipede in the family Geophilidae widespread in North America and named for the dark band of diamond-shaped markings across its back. G. vittatus grows up to 52 millimeters in length, though it averages 25–40, and ranges from brown- to orange-yellow in color, with 49–53 leg pairs in females, 49–51 leg pairs in males, and a lack of consolidated carpophagus structures. It can be found under any debris on the forest floor, but mostly under loose bark and occasionally near the sea.
Geophilus varians is a species of soil centipede in the family Geophilidae found in North America, particularly from South Carolina to Indiana, Ohio, Pennsylvania, and Virginia. It grows up to 40 millimeters, though it averages 30-35, ranges in color from light faded orange to yellow or whitish yellow with a deeper and brighter head, and has 53-59 leg pairs in males and 55-61 in females, as well as a complete lack of consolidated paxilli and sacculi, concealed prebasal plate, and unusually long ultimate legs.
Geophilus truncorum is a species of soil centipede in the family Geophilidae found across Western Europe, though it reaches as far as Poland, Italy, and Morocco. This centipede is relatively small, growing up to 20mm in length, with a yellow or orangeish brown body and dark yellow or brown head, denser and shorter hair than most Geophilus species, a main plate almost as elongated as in G. flavus (115:100), and distinct carpophagus fossae on the anterior sternites. Males of this species have 35 to 41 pairs of legs; females have 37 to 41.
Geophilus pusillifrater is a species of soil centipede in the family Geophilidae found in Bosnia-Herzegovina and Britain. It's a relatively small, pale species, prehensorial claws with pointed teeth at the base, distinct chitin lines, and a very broad ventral plate of the pregenital segment. It's at least partially a littoral creature. Males of this species have 39 or 41 pairs of legs; females have 43 leg pairs.
Geophilus pusillus is a species of soil centipede in the family Geophilidae found in Algeria. It grows up to 11 millimeters in length. The original description of this species is based on ten specimens: five male specimens from Algeria with 31 pairs of legs, and five specimens from Germany. Records from the Alpstein mountains indicate that G. pusillus is a soil-dwelling species that prefers humus-rich soil, but these records deserve confirmation. Some authorities consider the specimens recorded from Europe to be specimens of G. ribauti that have been misidentified as specimens of the Algerian species G. pusillus.
Geophilus punicus is a species of soil centipede in the family Geophilidae found in the Western Mediterranean, specifically Italy and North Africa (Tunisia). It's an epigeic species, though its habitat preferences are unknown apart from one record from caves, indicating it may also be a trogloxene. The original description of this species is based on male specimens with 43 pairs of legs.
Geophilus bobolianus is a species of soil centipede in the family Geophilidae found in France and Italy. This species has 45 to 51 pairs of legs. It was originally classified as a subspecies of G. longicornis identified by its lack of anterior sternal pores.
