Grullo

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A grulla, like all duns, exhibits a lighter body coat than mane and tail color, clear primitive markings (a distinctive dorsal stripe, horizontal striping on the back of the forelegs, often a transverse stripe over the withers), and the dark "dun mask" on the face. Zebra stripes are visible on the left back leg. The dun gene also produces light guard hairs in the mane and the tail. PoneyDun.jpg
A grulla, like all duns, exhibits a lighter body coat than mane and tail color, clear primitive markings (a distinctive dorsal stripe, horizontal striping on the back of the forelegs, often a transverse stripe over the withers), and the dark "dun mask" on the face. Zebra stripes are visible on the left back leg. The dun gene also produces light guard hairs in the mane and the tail.

Grulla, also called black dun, gray dun or mouse dun, is a color of horses in the dun family, characterized by tan-gray or mouse-colored hairs on the body, often with shoulder and dorsal stripes and black barring on the lower legs. The genotype for grulla horses is a black base with dun dilution. In this coloration, each individual hair is mouse-colored, unlike a roan, which is composed of a mixture of dark and light hairs. The several shades of grulla are informally referred to with a variety of terms, including black dun, blue dun, slate grulla, silver grulla or light grulla, silver dun, or lobo dun. Silver grulla may also refer to a grulla horse with silver dapple, regardless of shade. [1] In the Icelandic horse, the grulla color is called gray dun, in the Highland pony it is called mouse dun, and in the Norwegian Fjord horse, grå or gråblakk (literally, "gray dun").

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The word "grulla" in Spanish means crane. [2] (The original Spanish noun is pronounced [ˈɡɾuʝa] in American Spanish and [ˈɡɾuʎa] in Peninsular Spanish.)

A Heck horse Steppentarpan.jpg
A Heck horse

In terms of equine coat color genetics, all of these shades are based on the dun gene acting as a dilution gene over the black gene. Because the grulla color is not due to the gray gene, a grulla horse remains the same basic color from birth, though some minor shade variation may occur from summer to winter coats. If a grulla also carries the gray gene, it will be born a mouse tan-gray shade, usually with bold primitive markings, but then lighten and eventually develop a white hair coat with age. Because black is less common in general than bay or chestnut, grulla is likewise less common than red duns or bay (classic or zebra) duns. For example, only 0.7% of quarter horses registered each year with the AQHA are grulla.

The most obvious ways to tell whether a horse is grulla are not only the gray or tan-gray body color, but also its primitive markings, which include some or all of the following: dark face, cobwebbing around the eyes and forehead, dark mottling on the body, leg barring (sometimes called tiger striping), dark ear tips and edging, dark ear barring, dark shadowing of the neck, dark dorsal and transverse striping, and light guard hairs bordering a dark mane and tail.

Primitive roots

The tarpan (Equus ferus ferus) was a relative of the domestic horse that became extinct in the nineteenth century, and which appears to have had grulla coloration. The tarpan has been considered a true wild horse, an undomesticated relative or ancestor of the domestic horse. However, some authorities in the early twentieth century held the opinion that most equines called tarpans were actually domestic or feral horses, not a separate species. [3]

Several breeds with the grulla color have been developed in efforts to recreate ("breed back") the tarpan. These breeds include the Heck horse and Konik. One of the first experiments in this regard was published in 1906 by James Cossar Ewart, who obtained a "tarpan-like" horse by crossing a Shetland mare and a black Welsh pony. [3]

Related Research Articles

<span class="mw-page-title-main">Roan (color)</span>

Roan is a coat color found in many animals, including horses, cattle, antelope, cat and dogs. It is defined generally as an even mixture of white and pigmented hairs that do not "gray out" or fade as the animal ages. There are a variety of genetic conditions which produce the colors described as "roan" in various species.

<span class="mw-page-title-main">Palomino</span> Genetic color in horses

Palomino is a genetic color in horses, consisting of a gold coat and white mane and tail; the degree of whiteness can vary from bright white to yellow. Genetically, the palomino color is created by a single allele of a dilution gene called the cream gene working on a "red" (chestnut) base coat. Palomino is created by a genetic mechanism of incomplete dominance, hence it is not considered true-breeding. However, most color breed registries that record palomino horses were founded before equine coat color genetics were understood as well as they are today, therefore the standard definition of a palomino is based on the visible coat color, not heritability nor the underlying presence of the dilution gene.

<span class="mw-page-title-main">Tarpan</span> Extinct subspecies of equines

The term tarpan refers to free-ranging horses of the Eurasian steppe from the 18th to the 20th century. It is generally unknown whether those horses represented genuine wild horses, feral domestic horses or hybrids. The last individual believed to be a tarpan died in captivity in the Russian Empire in 1909.

<span class="mw-page-title-main">Bay (horse)</span> Hair coat color of horses

Bay is a hair coat color of horses, characterized by a reddish-brown or brown body color with a black point coloration on the mane, tail, ear edges, and lower legs. Bay is one of the most common coat colors in many horse breeds.

<span class="mw-page-title-main">Buckskin (horse)</span> Equine coat color

Buckskin is a breed of horse and is commonly misconceived for being merely the color of a horses coat. Buckskins coloring is a hair coat color referring to a color that resembles certain shades of tanned deerskin. Similar colors in some breeds of dogs are also called buckskin. The horse has a tan or gold colored coat with black points. Buckskin occurs as a result of the cream dilution gene acting on a bay horse. Therefore, a buckskin has the Extension, or "black base coat" (E) gene, the agouti gene (A) gene, which restricts the black base coat to the points, and one copy of the cream gene (CCr), which lightens the red/brown color of the bay coat to a tan/gold.

<span class="mw-page-title-main">Brindle</span> Coat coloring pattern in some animals

Brindle is a coat coloring pattern in animals, particularly dogs, cattle, guinea pigs, cats, and, rarely, horses. It is sometimes described as "tiger-striped", although the brindle pattern is more subtle than that of a tiger's coat.

<span class="mw-page-title-main">Point coloration</span> Coloration of animal coat/fur

Point coloration is animal coat coloration with a pale body and relatively darker extremities, i.e. the face, ears, feet, tail, and scrotum. It is most recognized as the coloration of Siamese and related breeds of cat, but can be found in dogs, rabbits, rats, sheep, guinea pigs and horses as well.

<span class="mw-page-title-main">Gray horse</span> Coat color characterized by progressive depigmentation of the colored hairs of the coat

A gray horse has a coat color characterized by progressive depigmentation of the colored hairs of the coat. Most gray horses have black skin and dark eyes; unlike some equine dilution genes and some other genes that lead to depigmentation, gray does not affect skin or eye color. Gray horses may be born any base color, depending on other color genes present. White hairs begin to appear at or shortly after birth and become progressively more prevalent as the horse ages as white hairs become intermingled with hairs of other colors. Graying can occur at different rates—very quickly on one horse and very slowly on another. As adults, most gray horses eventually become completely white, though some retain intermixed light and dark hairs.

<span class="mw-page-title-main">Cream gene</span> Gene for several horse coat colors

The cream gene is responsible for a number of horse coat colors. Horses that have the cream gene in addition to a base coat color that is chestnut will become palomino if they are heterozygous, having one copy of the cream gene, or cremello, if they are homozygous. Similarly, horses with a bay base coat and the cream gene will be buckskin or perlino. A black base coat with the cream gene becomes the not-always-recognized smoky black or a smoky cream. Cream horses, even those with blue eyes, are not white horses. Dilution coloring is also not related to any of the white spotting patterns.

<span class="mw-page-title-main">Champagne gene</span> Simple dominant allele responsible for a number of rare horse coat colors

The champagne gene is a simple dominant allele responsible for a number of rare horse coat colors. The most distinctive traits of horses with the champagne gene are the hazel eyes and pinkish, freckled skin, which are bright blue and bright pink at birth, respectively. The coat color is also affected: any hairs that would have been red are gold, and any hairs that would have been black are chocolate brown. If a horse inherits the champagne gene from either or both parents, a coat that would otherwise be chestnut is instead gold champagne, with bay corresponding to amber champagne, seal brown to sable champagne, and black to classic champagne. A horse must have at least one champagne parent to inherit the champagne gene, for which there is now a DNA test.

<span class="mw-page-title-main">Silver dapple gene</span>

The silver or silver dapple (Z) gene is a dilution gene that affects the black base coat color and is associated with Multiple Congenital Ocular Abnormalities. It will typically dilute a black mane and tail to a silvery gray or flaxen color, and a black body to a chocolaty brown, sometimes with dapples. It is responsible for a group of coat colors in horses called "silver dapple" in the west, or "taffy" in Australia. The most common colors in this category are black silver and bay silver, referring to the respective underlying coat color.

<span class="mw-page-title-main">Equine coat color genetics</span> Genetics behind the equine coat color

Equine coat color genetics determine a horse's coat color. Many colors are possible, but all variations are produced by changes in only a few genes. Bay is the most common color of horses. A change at the agouti locus is capable of turning bay to black, while a mutation at the extension locus can turn bay or black to chestnut.

<span class="mw-page-title-main">Dun gene</span> Dilution gene

The dun gene is a dilution gene that affects both red and black pigments in the coat color of a horse. The dun gene lightens most of the body while leaving the mane, tail, legs, and primitive markings the shade of the undiluted base coat color. A dun horse always has a dark dorsal stripe down the middle of its back, usually has a darker face and legs, and may have transverse striping across the shoulders or horizontal striping on the back of the forelegs. Body color depends on the underlying coat color genetics. A classic "bay dun" is a gray-gold or tan, characterized by a body color ranging from sandy yellow to reddish brown. Duns with a chestnut base may appear a light tan shade, and those with black base coloration are a smoky gray. Manes, tails, primitive markings, and other dark areas are usually the shade of the undiluted base coat color. The dun gene may interact with all other coat color alleles.

<span class="mw-page-title-main">Chestnut (horse color)</span> Horse coat color

Chestnut is a hair coat color of horses consisting of a reddish-to-brown coat with a mane and tail the same or lighter in color than the coat. Chestnut is characterized by the absolute absence of true black hairs. It is one of the most common horse coat colors, seen in almost every breed of horse.

<span class="mw-page-title-main">Equine coat color</span> Horse coat colors and markings

Horses exhibit a diverse array of coat colors and distinctive markings. A specialized vocabulary has evolved to describe them.

<span class="mw-page-title-main">Black horse</span> Horse coat color

Black is a hair coat color of horses in which the entire hair coat is black. Black is a relatively uncommon coat color, and it is not uncommon to mistake dark chestnuts or bays for black.

<span class="mw-page-title-main">Smoky black</span>

Smoky black or just smoky is a hair coat color of horses which appears dark brown to black in color. Smoky black is produced by the action of a heterozygous cream gene on an underlying black coat color. Therefore, smoky black is a member of the cream family of coat color dilutions, and found in horse populations that have other cream-based colors such as palomino, buckskin, perlino, cremello and smoky cream. All smoky blacks must have at least one parent with the cream gene, and a smoky black can only be verified through DNA testing or parentage. Smoky black has been mistaken for faded black, dark bay or brown, grullo or even liver chestnut.

<span class="mw-page-title-main">Primitive markings</span>

Primitive markings are a group of hair coat markings and qualities seen in several equine species, including horses, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such breeds. The markings are particularly associated with the dun coat color family. All dun horses possess at least the dorsal stripe, but the presence of the other primitive markings varies. Other common markings may include horizontal striping on the legs, transverse striping across the shoulders, and lighter guard hairs along the edges of a dark mane and tail.

<span class="mw-page-title-main">Seal brown (horse)</span> Hair coat color of horses

Seal brown is a hair coat color of horses characterized by a near-black body color; with black points, the mane, tail and legs; but also reddish or tan areas around the eyes, muzzle, behind the elbow and in front of the stifle. The term is not to be confused with "brown", which is used by some breed registries to refer to either a seal brown horse or to a dark bay without the additional characteristics of seal brown.

<span class="mw-page-title-main">Roan (horse)</span> Horse coat color pattern characterized by an even mixture of colored and white hairs on the body

Roan is a horse coat color pattern characterized by an even mixture of colored and white hairs on the body, while the head and "points"—lower legs, mane, and tail—are mostly solid-colored. Horses with roan coats have white hairs evenly intermingled throughout any other color. The head, legs, mane, and tail have fewer scattered white hairs or none at all. The roan pattern is dominantly inherited, and is found in many horse breeds. While the specific mutation responsible for roan has not been exactly identified, a DNA test can determine zygosity for roan in several breeds. True roan is always present at birth, though it may be hard to see until after the foal coat sheds out. The coat may lighten or darken from winter to summer, but unlike the gray coat color, which also begins with intermixed white and colored hairs, roans do not become progressively lighter in color as they age. The silvering effect of mixed white and colored hairs can create coats that look bluish or pinkish.

References

  1. "Grulla". Cedar Ridge Quarter Horses. Retrieved 2019-04-24.
  2. "Merriam-Webster Grulla" . Retrieved 2019-05-25.
  3. 1 2 J. C. Ewart (1906). "The tarpan and its relationship with wild and domestic horses". Nature. 74 (1909): 113–115. doi: 10.1038/074113a0 . [F]or more than a century all the horses living in a wild state in Europe, which happened to be of a mouse-dun color, seem to have been regarded as tarpans.

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